Increase in forest structural complexity along a precipitation gradient is mediated by partial harvests in temperate Patagonian forests.

Increasing forest structural complexity is becoming a common goal in forestry worldwide. However, the lack of empirical quantification clouds its implementation. Here we quantified the long-term effects (> 30 y) of partial harvest on stand structural complexity and net primary productivity using the east-west precipitation gradient (318-2508 mm, mean annual precipitation-MAP) of western Patagonian as a study system. In this gradient, pairs of 1-ha plots on 20 sites (20 plots harvested and 20 plots unharvested) were installed. In each plot terrestrial laser scanning was used to quantify the stand structural complexity index (SSCI), and Sentinel satellite images to obtain the Enhanced Vegetation Index (EVI: proxy of net primary productivity). Generalized linear mixed-effect models were used to relate SSCI to MAP and EVI to SSCI, with harvesting as indicator variable, and site as random variable (two plots nested to same precipitation). Results showed that harvested plots on mesic-to-humid sites (but not on dry sites) had higher SSCI and EVI values compared to unharvested plots, likely due to a greater vertical canopy packing. These results show the influence of precipitation on SSCI, which resulted in a more diversified stand structure and higher EVI. Such insights support site-specific management aimed to increase forest structural complexity.

Environmental factors affect the spatial arrangement of survival and damage of outplanted nothofagus dombeyi seedlings in the chilean andes / Factores ambientales afectan el arreglo espacial de la sobrevida y daño en plantas transplantadas de nothofagus dombeyi en los andes chilenos / Fatores ambientais afetam o arranjo espacial da sobrevida e dano em plantas transplantadas de nothofagus dombeyi nos andes chilenos

Mortality patterns were analyzed in a one-year old Nothofagus dombeyi plantation at mid-elevation in the Chilean Andes. Ripley´s univariate function was used to detect spatial patterns of mortality and damage (as reflected in crown dieback) of seedlings by assigning them into four categories: no crown damage, 1/3 of the crown damaged, 2/3 of the crown damaged and dead. Through correspondence analysis, variables (plant attributes, topography, weed competition, neighboring vegetation and fertilization) that could affect mortality were tested. At the end of the first growing season 67% of the seedlings survived, and by the end of the following dormant season only 37% were alive. Mortality patterns were random for seedlings with 1/3 of the crown damaged, and clustered for all other categories. Environmental variables with the greatest influence on mortality were increasing distance to a neighboring 10m tall plantation, absence of tall vegetation cover and convex micro-topography. Results suggest that large temperature oscillations with events of freezing temperatures (defined as the reported lethal temperature for 50% of its leaves) during the growing season, and severe frost during the dormant season, were the main causes of mortality and damage. The convenience of providing seedlings with some shelter when outplanted, or with an appropriate cold-acclimation treatment to resist low freezing temperatures when outplanted in open fields in harsh cold regions of the south-central Andes is discussed.

Se analizaron los patrones de mortandad en una plantación de Nothofagus dombeyi de un año de edad a altura media en los Andes chilenos. La función univariada de Ripley fue utilizada para detectar patrones espaciales de mortalidad y daño de las plantas asumiendo cuatro categorías: sin daño en la copa, 1/3 de copa dañada, 2/3 de copa dañada y muerte. Las variables (atributos de la planta, topografía, competencia de maleza, vegetación vecina y fertilización) fueron probadas por análisis de correspondencia. Al final de la primera estación de crecimiento 67% de las plantas sobrevivieron y al final del siguiente período latente solo 37% sobrevivían. Los patrones de mortalidad fueron aleatorios en plantas con 1/3 de la copa dañada, y agrupados en las otras tres categorías. Las variables ambientales con la mayor influencia en mortalidad fueron: distancia a una plantación vecina de 10m de altura, ausencia de cobertura vegetal alta y microtopografía convexa. Los resultados sugieren que grandes variaciones de temperatura con eventos de congelamiento (definido como la temperatura reportada como letal para 50% de las hojas) en la estación de crecimiento y congelamiento severo en la estación de latencia fueron las causas principales de mortalidad y daño. Se discute la conveniencia de proteger las plantaciones transplantadas o de una aclimatación apropiada para resistir las bajas temperaturas en plantas transplantadas a campo abierto en zonas frías de los Andes chilenos sur-centrales.

Analisaram-se os padrões de mortalidade em uma plantação de Nothofagus dombeyi de um ano de idade a altura média nos Andes chilenos. A função univariada de Ripley foi utilizada para detectar padrões espaciais de mortalidade e dano das plantas assumindo quatro categorias: sem dano na coroa, 1/3 de coroa danificada, 2/3 de coroa danificada e morte. As variáveis (atributos da planta, topografia, competência de maleza, vegetação vizinha e fertilização) foram provadas por análise de correspondência. No final da primeira estação de crescimento 67% das plantas sobreviveram e no final do seguinte período latente somente 37% sobreviviam. Os padrões de mortalidade foram aleatórios em plantas com 1/3 da coroa danificada, e agrupados nas outras três categorias. As variáveis ambientais com a maior influência em mortalidade foram: distância a uma plantação vizinha de 10m de altura, ausência de cobertura vegetal alta e microtopografía convexa. Os resultados sugerem que grandes variações de temperatura com momentos de congelamento (definido como a temperatura relatada como letal para 50% das folhas) na estação de crescimento e, congelamento severo na estação de latência, foram as causas principais de mortalidade e dano. Discute-se a conveniência de proteger as plantações transplantadas ou de uma aclimatação apropriada para resistir as baixas temperaturas em plantas transplantadas a campo aberto em zonas frias dos Andes chilenos sul-centrais.