Mapping and Analyzing Ecosystem Services Hotspots and Coldspots for Sustainable Spatial Planning in the Greater Asmara Area, Eritrea.

Rapid urbanization in African metropolises like the Greater Asmara Area, Eritrea, poses numerous environmental challenges, including soil sealing, loss of vegetation cover, threats to protected natural areas, and climate change, among others. Mapping and assessing ecosystem services, particularly analyzing their spatial and temporal distribution is crucial for sustainable spatial planning. This study aims at mapping and analyzing ecosystem services hotspots and coldspots dynamics in the Greater Asmara Area to identify recent trends and opportunities for enhancing ecosystem services supply. Utilizing remote sensing images, we produced land cover maps for 2009 and 2020 and mapped six ecosystem services through a lookup table approach. The study includes provisioning, regulating and maintenance, and cultural ecosystem services. We analyzed their spatio-temporal variations, identifying ecosystem services hotspots and coldspots and their changes over time. Results show that overall ecosystem services potential in the Greater Asmara Area remains low but stable, with some improvements. By 2020, areas with no ecosystem services potential decreased in southern regions like Gala Nefhi and Berik, and new hotspots and coldspots emerged in central Gala Nefhi. This pilot study demonstrates the feasibility and key challenges of the ecosystem services hotspots and coldspots approach for sustainable spatial planning in rapidly urbanizing African metropolitan regions. Despite limitations, the study offers valuable insights into ecosystem services potentials, and related hotspots and coldspots dynamics, raising awareness and paving the way for further research and application.

Simulating crowding of urban green areas to manage access during lockdowns.

During the COVID-19 emergency, cities around the world introduced measures to guarantee physical distancing that restricted access to urban parks and green areas, with potentially negative effects on citizens' health and wellbeing. This study aims at providing insights to manage access to urban green space in physical distancing times, when the risk of crowding should be avoided. Using the city of Trento (Italy) as a case study, the study simulates policy scenarios corresponding to different restrictions and assesses their effects on green space access and crowding. Policy scenarios are obtained by combining different distances that people are allowed to travel, different types of green areas available for public use (only urban parks or parks and schoolyards), and different target populations (all residents or only people with no private gardens). The results unveil the trade-off between access and crowding of green areas, and can be used to suggest policy interventions and regulations that can be adopted in an emergency. Particularly, the study shows that: i) The relationship between distance threshold and the percentage of people with access to green areas is non-linear, and this should be carefully considered when proposing travel restrictions; ii) Changing the maximum travel distance does not produce major effects on the number of crowded green areas, hence additional or alternative measures need to be adopted; iii) Off-the-shelf measures, such as opening schoolyards, are beneficial and can be implemented rapidly in an emergency. Finally, the study reveals "hotspots" of green space deprivation/overcrowding in the city that should be addressed by urban planning to ensure that green space continues to benefit citizens also during emergency conditions.

Governance Innovations for forest ecosystem service provision - Insights from an EU-wide survey.

This paper analyses the occurrence of governance innovations for forest ecosystem service (FES) provision in the forestry sector in Europe and the factors that influence innovation development. Based on a European-wide online survey, public and private forest owners and managers representing different property sizes indicate what type of governance innovation activities they engage in, and why. To investigate forestry innovations as systems, the analysis focuses on biophysical, social and technical factors influencing innovation development. The results of our exploratory quantitative analysis show that most innovation activities identified are largely oriented towards biomass production. Accordingly, most forest owners implement efficiency-driven optimisation strategies for forest management and technological improvement for provisioning service supply, to generate income. In contrast, the provision of regulating and cultural services is not yet a prominent part of forestry innovation activities.Reasons are rooted in a market-oriented economic rationale focusing on timber production, a lack of financial resources to compensate for other FES provisions or institutions to provide backup and security to forest owners and managers for engaging in innovation development. Given that the provision of a wide range of FES is a politically well-established objective for forest management in Europe, a strategy is needed that helps to align actors and sectors for supporting and co-financing related forest management approaches and business models. The current revision of the forest related policy framework on EU level under the EU Green deal poses a window of opportunity for better fostering novel governance approaches for more sustainable FES provision.

Multi-level climate change planning: An analysis of the Italian case.

As recognized by the Paris Climate Agreement of the United Nations Framework Convention on Climate Change (UNFCCC), local and subnational regions are crucial actors to achieve international mitigation and adaptation commitments. Scientific literature and empirical evidence point at multi-level climate governance as a crucial factor to engage subnational levels in the achievement of national and international objectives. This work focuses on the multi-level climate governance arrangements in Italy to investigate how Italian regions/provinces/cities are contributing to the achievement of national commitments. To this purpose, the paper undertakes a review of the climate policies of different tiers of government adopted to date and of the interrelationships among them. The results of the analysis show that the effective coordination between the different government levels should be strengthened to further incentivize and support initiatives at the local level. Results also show the relevant role played by international regional and city climate networks in boosting local and regional climate planning in Italy.

Climate mitigation in the Mediterranean Europe: An assessment of regional and city-level plans.

In Europe, regions in the Mediterranean area share common characteristics in terms of high sensitivity to climate change impacts. Does this translate into specificities regarding climate action that could arise from these Mediterranean characteristics? This paper sheds light on regional and local climate mitigation actions of the Mediterranean Europe, focusing on the plans to reduce greenhouse gases emissions in a representative sample of 51 regions and 73 cities across 9 Mediterranean countries (Croatia, Cyprus, France, Greece, Italy, Malta, Portugal, Slovenia, Spain). The study investigates: (i) the availability of local and regional mitigation plans, (ii) their goals in term of greenhouse gas emissions reduction targets on the short and medium-long term, and (iii) the impact of transnational climate networks on such local and regional climate mitigation planning. Results of this study indicate an uneven and fragmented planning, that shows a Mediterranean West-East divide, and a link with population size. However, overall, both regional and city action seem insufficiently ambitious with regards to meeting the Paris Agreement, at least at city level. While national frameworks are currently weak in influencing regional and local actions, transnational networks seem to be engaging factors for commitment (at city level) and ambitiousness (at regional level). The uneven and fragmented progress revealed by this study, does not align with the characteristics shared by investigated regions and cities in terms of environmental, socio-political, climatic and economic conditions. The results support the call of a common green deal at the Mediterranean level to further address specific Mediterranean challenges and related needs. This will allow to capitalise on available resources, generate local-specific knowledge, build capacities, and support Mediterranean regions and cities in preparing the next generation of more ambitious mitigation plans.

Assessing habitat quality in relation to the spatial distribution of protected areas in Italy.

The conservation of species and habitats is increasingly threatened by anthropogenic impacts, particularly land use change, from local to global scales. Although many efforts have been carried out so far to halt or at least reduce the biodiversity loss (e.g., the establishment of protected areas' networks), there are still both knowledge and policy gaps slowing the conservation of species and habitats in complex environments, such as the Mediterranean region. In particular, the human-driven impacts and threats on biodiversity need more careful analysis. Accordingly, this paper aims to assess the habitat quality and degradation in Italy in relation with the spatial pattern of the current protected areas' network, mainly to identify priority areas of intervention, thus supporting large-scale conservation strategies. A survey of experts was conducted to identify the main threats for biodiversity from different land uses at the national scale. The InVEST software was then applied to assess and map habitat quality and degradation with a high spatial resolution (20 m). The relationship between habitat quality and degradation as well as their hotspots, and alternative PA categories were also explored. Results indicate that: (i) habitat quality and degradation depend on the location and intensity of the anthropogenic impacts and are sensitive to different protection levels; (ii) the combination of the survey of experts and the spatially-explicit assessment of habitat quality and degradation is useful to highlight variations of the current conditions of biodiversity and habitats; and (iii) the identification of hotspots allows one to identify priority areas for conservation. Accordingly, the proposed approach may be used to strengthen the conservation efforts in similar contexts, and thus support the implementation of the biodiversity-related policies over the long term.

Ecosystem services classification: A systems ecology perspective of the cascade framework.

Ecosystem services research faces several challenges stemming from the plurality of interpretations of classifications and terminologies. In this paper we identify two main challenges with current ecosystem services classification systems: i) the inconsistency across concepts, terminology and definitions, and; ii) the mix up of processes and end-state benefits, or flows and assets. Although different ecosystem service definitions and interpretations can be valuable for enriching the research landscape, it is necessary to address the existing ambiguity to improve comparability among ecosystem-service-based approaches. Using the cascade framework as a reference, and Systems Ecology as a theoretical underpinning, we aim to address the ambiguity across typologies. The cascade framework links ecological processes with elements of human well-being following a pattern similar to a production chain. Systems Ecology is a long-established discipline which provides insight into complex relationships between people and the environment. We present a refreshed conceptualization of ecosystem services which can support ecosystem service assessment techniques and measurement. We combine the notions of biomass, information and interaction from system ecology, with the ecosystem services conceptualization to improve definitions and clarify terminology. We argue that ecosystem services should be defined as the interactions (i.e. processes) of the ecosystem that produce a change in human well-being, while ecosystem components or goods, i.e. countable as biomass units, are only proxies in the assessment of such changes. Furthermore, Systems Ecology can support a re-interpretation of the ecosystem services conceptualization and related applied research, where more emphasis is needed on the underpinning complexity of the ecological system.

Revitalizing traditional ecological knowledge: a study in an Alpine rural community.

This study aims to contribute to the debate on the value and the role of ecological knowledge in modern conservation strategies, with reference to the results of a case study conducted in the community of Montagne, located within a World Heritage site in the Italian Alps. This community is a paradigmatic example of the multiple transformations experienced by cultural landscapes in Alpine areas under the influence of global change. This study seeks to understand whether ecological knowledge is still in place in the community, and what the relationship is between the knowledge transmission and land use and social changes that have occurred in recent decades. To that end, the community is described by identifying the key variables (social, institutional, and ecological) that have historically shaped the landscape and the future priorities of the residents. Forest expansion, the most significant change in land use in the last 60 years, is analyzed using aerial photos; changes in biodiversity-related knowledge in the community are quantified by analyzing the inter-generational differences in plant species recognition. Results are discussed in the context of the current situation of the Montagne community, and the recommendation is made that policies and actions to promote traditional ecological knowledge protection or recovery in Europe be viewed as an important part of the recovery of community sovereignty and vitality. Lastly, concrete actions that can be implemented in our case study are proposed.

Geodesign to advance boundary work in urban planning: A study in Stockholm focused on nature-based solutions.

Geodesign supports collaborative urban planning by managing 'boundaries' between diverse knowledge holders. However, there is a paucity of empirical evidence of its contribution to 'boundary work'. This paper aims to evaluate how a geodesign process facilitates knowledge co-production through boundary work and to assess the scientific credibility, political saliency, and procedural legitimacy of its outputs in urban planning. We propose a replicable geodesign framework to assess boundary work, and test it in a case study on urban transformations with nature-based solutions in the Skarpnäck district of Stockholm, Sweden. Findings indicate that all geodesign steps facilitated communication by promoting collective reasoning. Participants acknowledged contributions to knowledge co-production and decision-making by mediating between different perspectives. However, data quality and modeling simplicity were identified as critical factors affecting the outputs' perceived credibility. Future applications should include co-designing the geodesign process, improving capacity and skills, and facilitating more integrated planning.

Understanding supply-demand mismatches in ecosystem services and interactive effects of drivers to support spatial planning in Tianjin metropolis, China.

Metropolitan areas are being challenged by the disparity between growing societal needs and dwindling natural resource provision. Understanding the supply-demand mismatches of ecosystem services (ES) and their drivers is essential for landscape planning and decision-making. However, integrating such information into spatial planning remains challenging due to the complex nature of urban ecosystems and their intrinsic interactions. In this study, we first assessed and mapped the supply, demand, and mismatches of six typical ES in Tianjin, China. We then clustered numerous townships based on their corresponding spatial characteristic of ES supply-demand mismatches. We also used Random Forest regression to examine the relative importance of drivers and applied Partial Least Squares structural equation modelling to decouple their interactions. The results showed that, the distribution of ES supply and demand showed obvious spatial heterogeneity, with a common surplus of ES supply in highly natural mountainous region and an excess of demand in urban centre. Additionally, all towns were classified into four spatial clusters with homogeneous states of supply-demand mismatches, serving as basic units for spatial optimization. Moreover, the interactions between drivers affected ES supply-demand mismatches in a coupled manner, including the direct effects of the socioeconomic factor (-0.821) and landscape composition (0.234), as well as the indirect effects of the biophysical factor (0.151) and landscape configuration (0.082). Finally, we discussed the utility of analysing the spatial mismatches between ES supply and demand for integrated territorial planning and coordinated decision-making.

Mapping and assessing ecosystem services for sustainable policy and decision-making in Eritrea.

The mapping and assessment of ecosystems and their services (MAES) is key to inform sustainable policy and decision-making at national and sub-national levels. Responding to the paucity of research in sub-Saharan Africa, we conduct a pilot study for Eritrea that aims to map and assess the temporal dynamics of key ecosystems and their services. We reviewed policy and legal documents, analyzed land cover changes and estimated the potential for ecosystem services supply through an expert-based matrix approach. Our results showed that from 2015 to 2019, the potential supply of the ecosystem services analyzed (e.g., crop provisioning, water supply and recreation) increased, with the exception of wood supply. Overall, our study presents policy-relevant insights as to where to conserve, develop, or restore ecosystem services supply in Eritrea. Our approach is transferable to similar data scarce contexts and can thereby support policies toward more sustainable land development for people and nature.

An environmental justice perspective on ecosystem services.

Mainstreaming of ecosystem service approaches has been proposed as one path toward sustainable development. Meanwhile, critics of ecosystem services question if the approach can account for the multiple values of ecosystems to diverse groups of people, or for aspects of inter- and intra-generational justice. In particular, an ecosystem service approach often overlooks power dimensions and capabilities that are core to environmental justice. This article addresses the need for greater guidance on incorporating justice into ecosystem services research and practice. We point to the importance of deep engagement with stakeholders and rights holders to disentangle contextual factors that moderate justice outcomes on ecosystem service attribution and appropriation in socio-political interventions. Such a holistic perspective enables the integration of values and knowledge plurality for enhancing justice in ecosystem services research. This broadened perspective paves a way for transformative ecosystem service assessments, management, and research, which can help inform and design governance structures that nourish human agency to sustainably identify, manage, and enjoy ecosystem services for human wellbeing.

Global shocks, cascading disruptions, and (re-)connections: viewing the COVID-19 pandemic as concurrent natural experiments to understand land system dynamics.

Context: For nearly three years, the COVID-19 pandemic has disrupted human well-being and livelihoods, communities, and economies in myriad ways with consequences for social-ecological systems across the planet. The pandemic represents a global shock in multiple dimensions that has already, and is likely to continue to have, far-reaching effects on land systems and on those depending on them for their livelihoods. Objectives: We focus on the observed effects of the pandemic on landscapes and people composing diverse land systems across the globe. Methods: We highlight the interrelated impacts of the pandemic shock on the economic, health, and mobility dimensions of land systems using six vignettes from different land systems on four continents, analyzed through the lens of socio-ecological resilience and the telecoupling framework. We present preliminary comparative insights gathered through interviews, surveys, key informants, and authors' observations and propose new research avenues for land system scientists. Results: The pandemic's effects have been unevenly distributed, context-specific, and dependent on the multiple connections that link land systems across the globe. Conclusions: We argue that the pandemic presents concurrent "natural experiments" that can advance our understanding of the intricate ways in which global shocks produce direct, indirect, and spillover effects on local and regional landscapes and land systems. These propagating shock effects disrupt existing connections, forge new connections, and re-establish former connections between peoples, landscapes, and land systems. Supplementary Information: The online version contains supplementary material available at 10.1007/s10980-023-01604-2.

Spatial rule-based assessment of habitat potential to predict impact of land use changes on biodiversity at municipal scale.

In human dominated landscapes, ecosystems are under increasing pressures caused by urbanization and infrastructure development. In Alpine valleys remnant natural areas are increasingly affected by habitat fragmentation and loss. In these contexts, there is a growing risk of local extinction for wildlife populations; hence assessing the consequences on biodiversity of proposed land use changes is extremely important. The article presents a methodology to assess the impacts of land use changes on target species at a local scale. The approach relies on the application of ecological profiles of target species for habitat potential (HP) assessment, using high resolution GIS-data within a multiple level framework. The HP, in this framework, is based on a species-specific assessment of the suitability of a site, as well of surrounding areas. This assessment is performed through spatial rules, structured as sets of queries on landscape objects. We show that by considering spatial dependencies in habitat assessment it is possible to perform better quantification of impacts of local-level land use changes on habitats.

Applying the ecosystem approach to select priority areas for forest landscape restoration in the Yungas, Northwestern Argentina.

This paper proposes a method to select forest restoration priority areas consistently with the key principles of the Ecosystem Approach (EA) and the Forest Landscape Restoration (FLR) framework. The methodology is based on the principles shared by the two approaches: acting at ecosystem scale, involving stakeholders, and evaluating alternatives. It proposes the involvement of social actors which have a stake in forest management through multicriteria analysis sessions aimed at identifying the most suitable forest restoration intervention. The method was applied to a study area in the native forests of Northern Argentina (the Yungas). Stakeholders were asked to identify alternative restoration actions, i.e. potential areas implementing FLR. Ten alternative fincas-estates derived from the Spanish land tenure system-differing in relation to ownership, management, land use, land tenure, and size were evaluated. Twenty criteria were selected and classified into four groups: biophysical, social, economic and political. Finca Ledesma was the closest to the economic, social, environmental and political goals, according to the values and views of the actors involved in the decision. This study represented the first attempt to apply EA principles to forest restoration at landscape scale in the Yungas region. The benefits obtained by the application of the method were twofold: on one hand, researchers and local actors were forced to conceive the Yungas as a complex net of rights rather than as a sum of personal interests. On the other hand, the participatory multicriteria approach provided a structured process for collective decision-making in an area where it has never been implemented.

Analysis of land suitability for the siting of inter-municipal landfills in the Cuitzeo Lake Basin, Mexico.

This paper presents three spatial decision-support models (Boolean logic, binary evidence and overlapping index of multiple class maps) to perform a land suitability analysis for sanitary landfill siting. The study was carried out in the basin of Lake Cuitzeo, Mexico, with the objective of locating areas that comply with environmental regulations and with the inter-municipality criterion, i.e., that are accessible by at least two municipalities. Biophysical and socio-economic data were processed in a Geographic Information System (GIS). The three models differ in their complexity and restrictiveness. The Boolean logic model is easier to apply and more restrictive than the other two, because it is based on the assessment of single attributes. On the other hand, the binary data and overlapping index methods are relatively more complex because they require attribute weighting. The results showed that 23 of the 28 municipalities included in the basin have at least one area that was classified as highly suitable. The most suitable areas covered from 63.8 to 204.5 km(2) (from 1.5% to 5%), and they are not distributed homogeneously, but clustered around four main sites. The larger and most suitable of these sites is located in the central part of the basin, and it can be accessed by five of the most densely populated municipalities. The proposed approach represents a low-cost alternative to support a common spatial decision-making process in developing countries.

The anthroposphere as an anticipatory system: Open questions on steering the climate.

Climate change research and action counteracting it affect everyone and would involve cross-societal transformations reshaping the anthroposphere in its entirety. Scrutinizing climate-related science and policies, we recognize attempts to steer the evolution of climate according to expected (or modelled) futures. Such attempts would turn the anthroposphere into a large "anticipatory system", in which human society seeks to anticipate and, possibly, to govern climate dynamics. The chief aim of this discussion paper is to open a critical debate on the climate change paradigm (CCP) drawing on a strategic and systemic framework grounded in the concept of anticipatory system sensu Rosen (1991). The proposed scheme is ambitiously intended to turn an intricate issue into a complex but structured problem that is to say, to make such complexity clear and manageable. This framework emerges from concepts borrowed from different scientific fields (including future studies and system dynamics) and its background lies in a simple quantitative literature overview, relying upon a broad level of analysis. The proposed framework will assist researchers and policy makers in thinking of CCP in terms of an anticipatory system, and in disentangling its interrelated (and sometimes intricate) aspects. In point of fact, several strategic questions related to CCP were not subjected to an adequate transdisciplinary discussion: what are the interplays between physical processes and social-political interventions, who is the observer (what he/she is looking for), and which paradigm is being used (or who defines the desirable future). The proposed scheme allows to structure such various topics in an arrangement which is easier to communicate, highlighting the linkages in between, and making them intelligible and open to verification and discussion. Furthermore, ideally developments will help scientists and policy makers address the strategic gaps between the evidence-based climatological assessments and the plurality of possible answers as applied to the geopolitical contingencies.

Boundary work for implementing adaptive management: A water sector application.

Boundary work, defined as effort to mediate between knowledge and action, is a promising approach for facilitating knowledge co-production for sustainable development. Here, we investigate a case study of knowledge co-production, to assess the applicability of boundary work as a conceptual framework to support implementing adaptive management in the water sector. We refer to a boundary work classification recently proposed by Clark et al., (2016), based on three types of knowledge uses, i.e. enlightenment, decision-, and negotiation-support, and three types of sources, i.e. personal expertise, single, and multiple communities of expertise. Our empirical results confirm boundary work has been crucial for the three types of knowledge use. For enlightenment and decision-support, effective interaction among knowledge producers and users was achieved through diverse boundary work practices, including joint agenda setting, and sharing of data and expertise. This initial boundary work eased subsequent knowledge co-production for decision-support and negotiations, in combination with stepping up of cooperation between relevant actors, suitable legislation and pressure for problem solving. Our analysis highlighted the temporal dimension matters - building trust around enlightenment first, and then using this as a basis for managing knowledge co-production for decision-, and negotiation support. We reconfirmed that boundary work is not a single time achievement, rather is a dynamic process, and we emphasized the importance of key actors driving the process, such as water utilities. Our results provide a rich case study of how strategic boundary work can facilitate knowledge co-production for adaptive management in the water sector. The boundary work practices employed here could also be transferred to other cases. Water utilities, as intermediaries between providers and beneficiaries of the important water-related ecosystem service of clean water provision, can indeed serve as key actors for initiating such boundary work practices.

Landscape changes, traditional ecological knowledge and future scenarios in the Alps: A holistic ecological approach.

In recent decades, a dramatic landscape change has occurred in the European alpine region: open areas have been naturally recolonized by forests as traditional agricultural and forest activities were reduced and reorganized. Land use changes (LUC) are generally measured through GIS and photo interpretation techniques, but despite many studies focused on this phenomenon and its effects on biodiversity and on the environment in general, there is a lack of information about the transformation of the human-environment connection. The study of Traditional Ecological Knowledge (TEK), such as the ability to recognize wild plants used as medicine or food, can suggest how this connection evolved through time and generations. This work investigates the relationship between the natural forest cover expansion that influences the loss of open areas and the loss of TEK. Different data sources and approaches were used to address the topic in all its complexity: a mix of questionnaire investigations, historical maps, GIS techniques and modelling were used to analyse past land use changes and predict future scenarios. The study area, Trentino, Italy, is paradigmatic of the alpine situation, and the land use change in the region is well documented by different studies, which were reviewed and compared in this paper. Our findings suggest that open area loss can be used as a good proxy to highlight the present state and to produce future scenarios of Traditional Ecological Knowledge. This could increase awareness of the loss of TEK in other Alpine regions, where data on TEK are lacking, but where environmental trends are comparable.

Modelling white-water rafting suitability in a hydropower regulated Alpine River.

Cultural and recreational river ecosystem services and their relations with the flow regime are still poorly investigated. We develop a modelling-based approach to assess recreational flow requirements and the spatially distributed river suitability for white-water rafting, a typical service offered by mountain streams, with potential conflicts of interest with hydropower regulation. The approach is based on the principles of habitat suitability modelling using water depth as the main attribute, with preference curves defined through interviews with local rafting guides. The methodology allows to compute streamflow thresholds for conditions of suitability and optimality of a river reach in relation to rafting. Rafting suitability response to past, present and future flow management scenarios can be predicted on the basis of a hydrological model, which is incorporated in the methodology and is able to account for anthropic effects. Rafting suitability is expressed through a novel metric, the "Rafting hydro-suitability index" (RHSI) which quantifies the cumulative duration of suitable and optimal conditions for rafting. The approach is applied on the Noce River (NE Italy), an Alpine River regulated by hydropower production and affected by hydropeaking, which influences suitability at a sub-daily scale. A dedicated algorithm is developed within the hydrological model to resemble hydropeaking conditions with daily flow data. In the Noce River, peak flows associated with hydropeaking support rafting activities in late summer, highlighting the dual nature of hydropeaking in regulated rivers. Rafting suitability is slightly reduced under present, hydropower-regulated flow conditions compared to an idealized flow regime characterised by no water abstractions. Localized water abstractions for small, run-of-the-river hydropower plants are predicted to negatively affect rafting suitability. The proposed methodology can be extended to support decision making for flow management in hydropower regulated streams, as it has the potential to quantify the response of different ecosystem services to flow regulation.