Asymmetric effects of sudden changes in timbre on auditory stream segregation.

Two experiments explored the effects of abrupt transitions in timbral properties [amplitude modulation (AM), pure tones vs narrow-band noises, and attack/decay envelope] on streaming. Listeners reported continuously the number of streams heard during 18-s-long alternating low- and high-frequency (LHL-) sequences (frequency separation: 2-6 semitones) that underwent a coherent transition at 6 s or remained unchanged. In experiment 1, triplets comprised unmodulated pure tones or 100%-depth AM was created using narrowly spaced tone pairs (dyads: 30- or 50-Hz modulation). In experiment 2, triplets comprised narrow-band noises, dyads, or pure tones with quasi-trapezoidal envelopes (10/80/10 ms), fast attacks and slow decays (10/90 ms), or vice versa (90/10 ms). Abrupt transitions led to direction-dependent changes in stream segregation. Transitions from modulated to unmodulated (or slower-modulated) tones, from noise bands to pure tones, or from slow- to fast-attack tones typically caused substantial loss of segregation (resetting), whereas transitions in the opposite direction mostly caused less or no resetting. Furthermore, for the smallest frequency separation, transitions in the latter direction usually led to increased segregation (overshoot). Overall, the results are reminiscent of the perceptual asymmetries found in auditory search for targets with or without a salient additional feature (or greater activation of that feature).

The influence of envelope shape on the lateralization of amplitude-modulated, low-frequency sound.

For abruptly gated sound, interaural time difference (ITD) cues at onset carry greater perceptual weight than those following. This research explored how envelope shape influences such carrier ITD weighting. Experiment 1 assessed the perceived lateralization of a tonal binaural beat that transitioned through ITD (diotic envelope, mean carrier frequency of 500 Hz). Listeners' left/right lateralization judgments were compared to those for static-ITD tones. For an 8 Hz sinusoidally amplitude-modulated envelope, ITD cues 24 ms after onset well-predicted reported sidedness. For an equivalent-duration "abrupt" envelope, which was unmodulated besides 20-ms onset/offset ramps, reported sidedness corresponded to ITDs near onset (e.g., 6 ms). However, unlike for sinusoidal amplitude modulation, ITDs toward offset seemingly also influenced perceived sidedness. Experiment 2 adjusted the duration of the offset ramp (25-75 ms) and found evidence for such offset weighting only for the most abrupt ramp tested. In experiment 3, an ITD was imposed on a brief segment of otherwise diotic filtered noise. Listeners discriminated right- from left-leading ITDs. In sinusoidal amplitude modulation, thresholds were lowest when the ITD segment occurred during rising amplitude. For the abrupt envelope, the lowest thresholds were observed when the segment occurred at either onset or offset. These experiments demonstrate the influence of envelope profile on carrier ITD sensitivity.

Estimating the perceptual weighting of interaural time difference cues in amplitude modulated binaural beats.

For an abruptly gated sound, perceived lateralization is determined primarily by binaural cues at onset. Relatively less is known about the temporal weighing of binaural cues-such as interaural time difference (ITD)-during more naturalistic modulation profiles. Here, an experiment measured the lateralization of a tonal binaural beat modulated by a diotic, 8-Hz sinusoidal amplitude modulation. Binaural beat lateralization (left/right, two alternatives) was compared to that for tones with static ITDs. Across three mean carrier frequencies (200, 500, and 800 Hz), ITDs occurring during early rising amplitude (e.g., 20-25 ms after onset) predicted the perceived lateralization of the binaural beat signals well.

Effects of inducer continuity on auditory stream segregation: comparison of physical and perceived continuity in different contexts.

The factors influencing the stream segregation of discrete tones and the perceived continuity of discrete tones as continuing through an interrupting masker are well understood as separate phenomena. Two experiments tested whether perceived continuity can influence the build-up of stream segregation by manipulating the perception of continuity during an induction sequence and measuring streaming in a subsequent test sequence comprising three triplets of low and high frequency tones (LHL-[ellipsis (horizontal)]). For experiment 1, a 1.2-s standard induction sequence comprising six 100-ms L-tones strongly promoted segregation, whereas a single extended L-inducer (1.1 s plus 100-ms silence) did not. Segregation was similar to that following the single extended inducer when perceived continuity was evoked by inserting noise bursts between the individual tones. Reported segregation increased when the noise level was reduced such that perceived continuity no longer occurred. Experiment 2 presented a 1.3-s continuous inducer created by bridging the 100-ms silence between an extended L-inducer and the first test-sequence tone. This configuration strongly promoted segregation. Segregation was also increased by filling the silence after the extended inducer with noise, such that it was perceived like a bridging inducer. Like physical continuity, perceived continuity can promote or reduce test-sequence streaming, depending on stimulus context.

Perceived tonal continuity through two noise bursts separated by silence.

Three experiments measured the perceived continuity of two pure tones "flankers" through a masker containing a silence. Experiment 1 used a 2I-2AFC procedure; one interval contained two noise bursts separated by a silent gap, and the other contained two noise bursts separated by a tone of the same duration as the silence. Discrimination between masker conditions was very accurate when the flankers were absent but was impaired substantially when the flankers were present. This was taken as evidence that illusory flanker continuity during the silent gap was heard as similar to the physical presence of a tone in the gap. In experiment 2, performance remained poor when the flankers were frequency glides aligned along a common trajectory. Performance improved significantly when the flankers were misaligned in trajectory. In experiment 3, listeners rated directly perceived flanker continuity. Strong continuity was reported in the silent gap conditions for which poor performance had been observed in experiments 1 and 2. These findings show that continuity may be heard through a masker that cannot mask a physically continuous tone but can mask the flankers' offset and onset. The results are explained in terms of the perceptual grouping of onsets and offsets of the flankers.

Build-up of the tendency to segregate auditory streams: resetting effects evoked by a single deviant tone.

The tendency to hear a sequence of alternating low (L) and high (H) frequency tones as two streams can be increased by a preceding induction sequence, even one composed only of same-frequency tones. Four experiments used such an induction sequence (10 identical L tones) to promote segregation in a shorter test sequence comprising L and H tones. Previous studies have shown that the build-up of stream segregation is usually reduced greatly when a sudden change in acoustic properties distinguishes all of the induction tones from their test-sequence counterparts. Experiment 1 showed that a single deviant tone, created by altering the final inducer (in frequency, level, duration, or replacement with silence) reduced reported segregation, often substantially. Experiment 2 partially replicated this finding, using changes in temporal discrimination as a measure of streaming. Experiments 3 and 4 varied the size of a frequency change applied to the deviant tone; the extent of resetting varied with size only gradually. The results suggest that resetting begins to occur once the change is large enough to be noticeable. Since the prior inducers always remained unaltered in the deviant-tone conditions, it is proposed that a single change actively resets the build-up evoked by the induction sequence.

A Comparison of Two Objective Measures of Binaural Processing: The Interaural Phase Modulation Following Response and the Binaural Interaction Component.

There has been continued interest in clinical objective measures of binaural processing. One commonly proposed measure is the binaural interaction component (BIC), which is obtained typically by recording auditory brainstem responses (ABRs)-the BIC reflects the difference between the binaural ABR and the sum of the monaural ABRs (i.e., binaural - (left + right)). We have recently developed an alternative, direct measure of sensitivity to interaural time differences, namely, a following response to modulations in interaural phase difference (the interaural phase modulation following response; IPM-FR). To obtain this measure, an ongoing diotically amplitude-modulated signal is presented, and the interaural phase difference of the carrier is switched periodically at minima in the modulation cycle. Such periodic modulations to interaural phase difference can evoke a steady state following response. BIC and IPM-FR measurements were compared from 10 normal-hearing subjects using a 16-channel electroencephalographic system. Both ABRs and IPM-FRs were observed most clearly from similar electrode locations-differential recordings taken from electrodes near the ear (e.g., mastoid) in reference to a vertex electrode (Cz). Although all subjects displayed clear ABRs, the BIC was not reliably observed. In contrast, the IPM-FR typically elicited a robust and significant response. In addition, the IPM-FR measure required a considerably shorter recording session. As the IPM-FR magnitude varied with interaural phase difference modulation depth, it could potentially serve as a correlate of perceptual salience. Overall, the IPM-FR appears a more suitable clinical measure than the BIC.

Build-up of auditory stream segregation induced by tone sequences of constant or alternating frequency and the resetting effects of single deviants.

A sequence of constant-frequency tones can promote streaming in a subsequent sequence of alternating-frequency tones, but why this effect occurs is not fully understood and its time course has not been investigated. Experiment 1 used a 2.0-s-long constant-frequency inducer (10 repetitions of a low-frequency pure tone) to promote segregation in a subsequent, 1.2-s test sequence of alternating low- and high-frequency tones. Replacing the final inducer tone with silence substantially reduced reported test-sequence segregation. This reduction did not occur when either the 4th or 7th inducer was replaced with silence. This suggests that a change at the induction/test-sequence boundary actively resets build-up, rather than less segregation occurring simply because fewer inducer tones were presented. Furthermore, Experiment 2 found that a constant-frequency inducer produced its maximum segregation-promoting effect after only three tones--this contrasts with the more gradual build-up typically observed for alternating-frequency sequences. Experiment 3 required listeners to judge continuously the grouping of 20-s test sequences. Constant-frequency inducers were considerably more effective at promoting segregation than alternating ones; this difference persisted for ∼10 s. In addition, resetting arising from a single deviant (longer tone) was associated only with constant-frequency inducers. Overall, the results suggest that constant-frequency inducers promote segregation by capturing one subset of test-sequence tones into an ongoing, preestablished stream, and that a deviant tone may reduce segregation by disrupting this capture. These findings offer new insight into the dynamics of stream segregation, and have implications for the neural basis of streaming and the role of attention in stream formation.

Sequential grouping of pure-tone percepts evoked by the segregation of components from a complex tone.

A sudden change applied to a single component can cause its segregation from an ongoing complex tone as a pure-tone-like percept. Three experiments examined whether such pure-tone-like percepts are organized into streams by extending the research of Bregman and Rudnicky (1975). Those authors found that listeners struggled to identify the presentation order of 2 pure-tone targets of different frequency when they were flanked by 2 lower frequency "distractors." Adding a series of matched-frequency "captor" tones, however, improved performance by pulling the distractors into a separate stream from the targets. In the current study, sequences of discrete pure tones were substituted by sequences of brief changes applied to an otherwise constant 1.2-s complex tone. Pure-tone-like percepts were evoked by applying 6-dB increments to individual components of a complex comprising harmonics 1-7 of 300 Hz (Experiment 1) or 0.5-ms changes in interaural time difference to individual components of a log-spaced complex (range 160-905 Hz; Experiment 2). Results were consistent with the earlier study, providing clear evidence that pure-tone-like percepts are organized into streams. Experiment 3 adapted Experiment 1 by presenting a global amplitude increment either synchronous with, or just after, the last captor prior to the 1st distractor. In the former case, for which there was no pure-tone-like percept corresponding to that captor, the captor sequence did not aid performance to the same extent as previously. It is concluded that this change to the captor-tone stream partially resets the stream-formation process, and so the distractors and targets became likely to integrate once more.