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1.
Proc Natl Acad Sci U S A ; 119(42): e2214005119, 2022 10 18.
Artigo em Inglês | MEDLINE | ID: mdl-36215511

RESUMO

How does the mind make moral judgments when the only way to satisfy one moral value is to neglect another? Moral dilemmas posed a recurrent adaptive problem for ancestral hominins, whose cooperative social life created multiple responsibilities to others. For many dilemmas, striking a balance between two conflicting values (a compromise judgment) would have promoted fitness better than neglecting one value to fully satisfy the other (an extreme judgment). We propose that natural selection favored the evolution of a cognitive system designed for making trade-offs between conflicting moral values. Its nonconscious computations respond to dilemmas by constructing "rightness functions": temporary representations specific to the situation at hand. A rightness function represents, in compact form, an ordering of all the solutions that the mind can conceive of (whether feasible or not) in terms of moral rightness. An optimizing algorithm selects, among the feasible solutions, one with the highest level of rightness. The moral trade-off system hypothesis makes various novel predictions: People make compromise judgments, judgments respond to incentives, judgments respect the axioms of rational choice, and judgments respond coherently to morally relevant variables (such as willingness, fairness, and reciprocity). We successfully tested these predictions using a new trolley-like dilemma. This dilemma has two original features: It admits both extreme and compromise judgments, and it allows incentives-in this case, the human cost of saving lives-to be varied systematically. No other existing model predicts the experimental results, which contradict an influential dual-process model.


Assuntos
Julgamento , Princípios Morais , Humanos , Motivação , Comportamento Social
2.
Behav Brain Sci ; 46: e300, 2023 10 04.
Artigo em Inglês | MEDLINE | ID: mdl-37789553

RESUMO

Self-control provides one cooperative explanation for "purity." Other types of cooperation provide additional explanations. For example, individuals compete for status by displaying high-value social and sexual traits, which are moralised because they reduce the mutual costs of conflict. As this theory predicts, sexually unattractive traits are perceived as morally bad, aside from self-control. Moral psychology will advance more quickly by drawing on all theories of cooperation.


Assuntos
Comportamento Cooperativo , Princípios Morais , Humanos , Comportamento Sexual
4.
Proc Natl Acad Sci U S A ; 115(33): 8322-8327, 2018 08 14.
Artigo em Inglês | MEDLINE | ID: mdl-30068602

RESUMO

Becoming valuable to fellow group members so that one would attract assistance in times of need is a major adaptive problem. To solve it, the individual needs a predictive map of the degree to which others value different acts so that, in choosing how to act, the payoff arising from others' valuation of a potential action (e.g., showing bandmates that one is a skilled forager by pursuing a hard-to-acquire prey item) can be added to the direct payoff of the action (e.g., gaining the nutrients of the prey captured). The pride system seems to incorporate all of the elements necessary to solve this adaptive problem. Importantly, data from western(-ized), educated, industrialized, rich, and democratic (WEIRD) societies indicate close quantitative correspondences between pride and the valuations of audiences. Do those results generalize beyond industrial mass societies? To find out, we conducted an experiment among 567 participants in 10 small-scale societies scattered across Central and South America, Africa, and Asia: (i) Bosawás Reserve, Nicaragua; (ii) Cotopaxi, Ecuador; (iii) Drâa-Tafilalet, Morocco; (iv) Enugu, Nigeria; (v) Le Morne, Mauritius; (vi) La Gaulette, Mauritius; (vii) Tuva, Russia; (viii) Shaanxi and Henan, China; (ix) farming communities in Japan; and (x) fishing communities in Japan. Despite widely varying languages, cultures, and subsistence modes, pride in each community closely tracked the valuation of audiences locally (mean r = +0.66) and even across communities (mean r = +0.29). This suggests that the pride system not only develops the same functional architecture everywhere but also operates with a substantial degree of universality in its content.


Assuntos
Cognição , Emoções , Comportamento Social , Comparação Transcultural , Humanos , Sociedades
5.
Proc Natl Acad Sci U S A ; 115(39): 9702-9707, 2018 09 25.
Artigo em Inglês | MEDLINE | ID: mdl-30201711

RESUMO

Human foragers are obligately group-living, and their high dependence on mutual aid is believed to have characterized our species' social evolution. It was therefore a central adaptive problem for our ancestors to avoid damaging the willingness of other group members to render them assistance. Cognitively, this requires a predictive map of the degree to which others would devalue the individual based on each of various possible acts. With such a map, an individual can avoid socially costly behaviors by anticipating how much audience devaluation a potential action (e.g., stealing) would cause and weigh this against the action's direct payoff (e.g., acquiring). The shame system manifests all of the functional properties required to solve this adaptive problem, with the aversive intensity of shame encoding the social cost. Previous data from three Western(ized) societies indicated that the shame evoked when the individual anticipates committing various acts closely tracks the magnitude of devaluation expressed by audiences in response to those acts. Here we report data supporting the broader claim that shame is a basic part of human biology. We conducted an experiment among 899 participants in 15 small-scale communities scattered around the world. Despite widely varying languages, cultures, and subsistence modes, shame in each community closely tracked the devaluation of local audiences (mean r = +0.84). The fact that the same pattern is encountered in such mutually remote communities suggests that shame's match to audience devaluation is a design feature crafted by selection and not a product of cultural contact or convergent cultural evolution.


Assuntos
Comparação Transcultural , Vergonha , Cultura , Feminino , Humanos , Masculino , Características de Residência , Comportamento Social
6.
Proc Natl Acad Sci U S A ; 114(31): 8420-8425, 2017 08 01.
Artigo em Inglês | MEDLINE | ID: mdl-28716928

RESUMO

Why do people support economic redistribution? Hypotheses include inequity aversion, a moral sense that inequality is intrinsically unfair, and cultural explanations such as exposure to and assimilation of culturally transmitted ideologies. However, humans have been interacting with worse-off and better-off individuals over evolutionary time, and our motivational systems may have been naturally selected to navigate the opportunities and challenges posed by such recurrent interactions. We hypothesize that modern redistribution is perceived as an ancestral scene involving three notional players: the needy other, the better-off other, and the actor herself. We explore how three motivational systems-compassion, self-interest, and envy-guide responses to the needy other and the better-off other, and how they pattern responses to redistribution. Data from the United States, the United Kingdom, India, and Israel support this model. Endorsement of redistribution is independently predicted by dispositional compassion, dispositional envy, and the expectation of personal gain from redistribution. By contrast, a taste for fairness, in the sense of (i) universality in the application of laws and standards, or (ii) low variance in group-level payoffs, fails to predict attitudes about redistribution.


Assuntos
Empatia/fisiologia , Motivação/fisiologia , Comportamento Social , Seguridade Social/psicologia , Atitude , Feminino , Humanos , Índia , Israel , Masculino , Princípios Morais , Fatores Socioeconômicos , Reino Unido , Estados Unidos
7.
Proc Natl Acad Sci U S A ; 114(8): 1874-1879, 2017 02 21.
Artigo em Inglês | MEDLINE | ID: mdl-28167752

RESUMO

Pride occurs in every known culture, appears early in development, is reliably triggered by achievements and formidability, and causes a characteristic display that is recognized everywhere. Here, we evaluate the theory that pride evolved to guide decisions relevant to pursuing actions that enhance valuation and respect for a person in the minds of others. By hypothesis, pride is a neurocomputational program tailored by selection to orchestrate cognition and behavior in the service of: (i) motivating the cost-effective pursuit of courses of action that would increase others' valuations and respect of the individual, (ii) motivating the advertisement of acts or characteristics whose recognition by others would lead them to enhance their evaluations of the individual, and (iii) mobilizing the individual to take advantage of the resulting enhanced social landscape. To modulate how much to invest in actions that might lead to enhanced evaluations by others, the pride system must forecast the magnitude of the evaluations the action would evoke in the audience and calibrate its activation proportionally. We tested this prediction in 16 countries across 4 continents (n = 2,085), for 25 acts and traits. As predicted, the pride intensity for a given act or trait closely tracks the valuations of audiences, local (mean r = +0.82) and foreign (mean r = +0.75). This relationship is specific to pride and does not generalize to other positive emotions that coactivate with pride but lack its audience-recalibrating function.


Assuntos
Cognição , Comparação Transcultural , Emoções , Comportamento Social , Comportamento de Escolha , Feminino , Humanos , Masculino , Motivação
8.
Proc Natl Acad Sci U S A ; 113(10): 2625-30, 2016 Mar 08.
Artigo em Inglês | MEDLINE | ID: mdl-26903649

RESUMO

We test the theory that shame evolved as a defense against being devalued by others. By hypothesis, shame is a neurocomputational program tailored by selection to orchestrate cognition, motivation, physiology, and behavior in the service of: (i) deterring the individual from making choices where the prospective costs of devaluation exceed the benefits, (ii) preventing negative information about the self from reaching others, and (iii) minimizing the adverse effects of devaluation when it occurs. Because the unnecessary activation of a defense is costly, the shame system should estimate the magnitude of the devaluative threat and use those estimates to cost-effectively calibrate its activation: Traits or actions that elicit more negative evaluations from others should elicit more shame. As predicted, shame closely tracks the threat of devaluation in the United States (r = .69), India (r = .79), and Israel (r = .67). Moreover, shame in each country strongly tracks devaluation in the others, suggesting that shame and devaluation are informed by a common species-wide logic of social valuation. The shame-devaluation link is also specific: Sadness and anxiety-emotions that coactivate with shame-fail to track devaluation. To our knowledge, this constitutes the first empirical demonstration of a close, specific match between shame and devaluation within and across cultures.


Assuntos
Cognição/fisiologia , Motivação/fisiologia , Autoimagem , Vergonha , Comparação Transcultural , Emoções/fisiologia , Humanos , Índia , Israel , Modelos Psicológicos , Psicologia do Self , Percepção Social , Estresse Psicológico/psicologia , Estados Unidos
9.
Behav Brain Sci ; 41: e189, 2018 01.
Artigo em Inglês | MEDLINE | ID: mdl-31064534

RESUMO

Some questions, such as when a statistical distribution of incomes becomes too unequal, seem highly attention-grabbing, inferentially productive, and morally vexing. Yet many other questions that are crucial to the functioning of a modern economy seem uninteresting non-issues. An evolutionary-psychological framework to study folk-economic beliefs has the potential to illuminate this puzzle.


Assuntos
Evolução Biológica , Eficiência , Cognição , Fatores Socioeconômicos
10.
Evol Hum Behav ; 36(1): 44-51, 2015 Jan 01.
Artigo em Inglês | MEDLINE | ID: mdl-25663798

RESUMO

Prior to, or concurrent with, the encoding of concepts into speech, the individual faces decisions about whether, what, when, how, and with whom to communicate. Compared to the existing wealth of linguistic knowledge however, we know little of the mechanisms that govern the delivery and accrual of information. Here we focus on a fundamental issue of communication: The decision whether to deliver information. Specifically, we study spontaneous confession to a victim. Given the costs of social devaluation, offenders are hypothesized to refrain from confessing unless the expected benefits of confession (e.g. enabling the victim to remedially modify their course of action) outweigh its marginal costs-the victim's reaction, discounted by the likelihood that information about the offense has not leaked. The logic of welfare tradeoffs indicates that the victim's reaction will be less severe and, therefore, less costly to the offender, with decreases in the cost of the offense to the victim and, counter-intuitively, with increases in the benefit of the offense to the offender. Data from naturalistic offenses and experimental studies supported these predictions. Offenders are more willing to confess when the benefit of the offense to them is high, the cost to the victim is low, and the probability of information leakage is high. This suggests a conflict of interests between senders and receivers: Often, offenders are more willing to confess when confessions are less beneficial to the victims. An evolutionary-computational framework is a fruitful approach to understanding the factors that regulate communication.

11.
Emotion ; 24(5): 1157-1168, 2024 Aug.
Artigo em Inglês | MEDLINE | ID: mdl-38252113

RESUMO

The emotions of guilt and shame play major roles in forgiveness, social exclusion, face-saving ploys, suicide, and honor killings. Understanding these emotions is thus of vital importance. The outputs of guilt and shame are already well understood: Guilt motivates amends; shame motivates evasion. However, the elicitors and functions of these emotions are disputed. According to attributional theory, guilt and shame are intrapersonal emotions elicited when negative outcomes are attributed to controllable/unstable (guilt) or uncontrollable/stable (shame) aspects of the self. By contrast, functionalist theory claims that guilt and shame are interpersonal emotions for minimizing the imposition of harm on valued others (guilt) and the cost of reputational damage on the self (shame). Although there is confirmatory evidence consistent with both theories, evidence ostensibly supporting one theory has been argued to actually support the other. To solve this problem of data interpretation, here we report contrastive critical tests of the two theories performed on online participant pools in the United States and India in 2021 (N = 853). Results in both countries support functionalist theory over attributional theory, suggesting that the intrapersonal effects reported in the emotion literature are tributary or incidental to the interpersonal functions of guilt and shame. Functionalist theory presents a promising framework for understanding the interpersonal and intrapersonal aspects of guilt, shame, and other self-conscious emotions. (PsycInfo Database Record (c) 2024 APA, all rights reserved).


Assuntos
Culpa , Autoimagem , Vergonha , Humanos , Feminino , Masculino , Adulto , Estados Unidos , Índia , Teoria Psicológica , Comparação Transcultural , Adulto Jovem , Emoções/fisiologia , Relações Interpessoais
12.
Psychol Sci ; 24(7): 1098-103, 2013 Jul 01.
Artigo em Inglês | MEDLINE | ID: mdl-23670886

RESUMO

Over human evolutionary history, upper-body strength has been a major component of fighting ability. Evolutionary models of animal conflict predict that actors with greater fighting ability will more actively attempt to acquire or defend resources than less formidable contestants will. Here, we applied these models to political decision making about redistribution of income and wealth among modern humans. In studies conducted in Argentina, Denmark, and the United States, men with greater upper-body strength more strongly endorsed the self-beneficial position: Among men of lower socioeconomic status (SES), strength predicted increased support for redistribution; among men of higher SES, strength predicted increased opposition to redistribution. Because personal upper-body strength is irrelevant to payoffs from economic policies in modern mass democracies, the continuing role of strength suggests that modern political decision making is shaped by an evolved psychology designed for small-scale groups.


Assuntos
Tomada de Decisões , Economia , Renda , Força Muscular , Política , Comportamento Social , Adolescente , Adulto , Agressão , Argentina , Braço/anatomia & histologia , Dinamarca , Feminino , Humanos , Masculino , Pessoa de Meia-Idade , Contração Muscular , Músculo Esquelético/anatomia & histologia , Classe Social , Estados Unidos , Adulto Jovem
13.
Evol Psychol ; 21(3): 14747049231203394, 2023.
Artigo em Inglês | MEDLINE | ID: mdl-37770021

RESUMO

Previous research indicates that the anticipatory shame an individual feels at the prospect of taking a disgraceful action closely tracks the degree to which local audiences, and even foreign audiences, devalue those individuals who take that action. This supports the proposition that the shame system (a) defends the individual against the threat of being devalued, and (b) balances the competing demands of operating effectively yet efficiently. The stimuli events used in previous research were highly variable in their perceived disgracefulness, ranging in rated shame and audience devaluation from low (e.g., missing the target in a throwing game) to high (e.g., being discovered cheating on one's spouse). But how precise is the tracking of audience devaluation by the shame system? Would shame track devaluation for events that are similarly low (or high) in disgracefulness? To answer this question, we conducted a study with participants from the United States and India. Participants were assigned, between-subjects, to one of two conditions: shame or audience devaluation. Within-subjects, participants rated three low-variation sets of 25 scenarios each, adapted from Mu, Kitayama, Han, & Gelfand (2015), which convey (a) appropriateness (e.g., yelling at a rock concert), (b) mild disgracefulness (e.g., yelling on the metro), and (c) disgracefulness (e.g., yelling in the library), all presented un-blocked, in random order. Consistent with previous research, shame tracked audience devaluation across the high-variation superset of 75 scenarios, both within and between cultures. Critically, shame tracked devaluation also within each of the three sets. The shame system operates with high precision.


Assuntos
Emoções , Vergonha , Humanos , Estados Unidos , Processos Mentais
14.
Evol Hum Sci ; 4: e45, 2022.
Artigo em Inglês | MEDLINE | ID: mdl-37588893

RESUMO

The shame system appears to be natural selection's solution to the adaptive problem of information-triggered reputational damage. Over evolutionary time, this problem would have led to a coordinated set of adaptations - the shame system - designed to minimise the spread of negative information about the self and the likelihood and costs of being socially devalued by others. This information threat theory of shame can account for much of what we know about shame and generate precise predictions. Here, we analyse the behavioural configuration that people adopt stereotypically when ashamed - slumped posture, downward head tilt, gaze avoidance, inhibition of speech - in light of shame's hypothesised function. This behavioural configuration may have differentially favoured its own replication by (a) hampering the transfer of information (e.g. diminishing audiences' tendency to attend to or encode identifying information - shame camouflage) and/or (b) evoking less severe devaluative responses from audiences (shame display). The shame display hypothesis has received considerable attention and empirical support, whereas the shame camouflage hypothesis has to our knowledge not been advanced or tested. We elaborate on this hypothesis and suggest directions for future research on the shame pose.

15.
Evol Hum Sci ; 3: e10, 2021.
Artigo em Inglês | MEDLINE | ID: mdl-37588558

RESUMO

The emotion of pride appears to be a neurocognitive guidance system to capitalize on opportunities to become more highly valued and respected by others. Whereas the inputs and the outputs of pride are relatively well understood, little is known about how the pride system matches inputs to outputs. How does pride work? Here we evaluate the hypothesis that pride magnitude matches the various outputs it controls to the present activating conditions - the precise degree to which others would value the focal individual if the individual achieved a particular achievement. Operating in this manner would allow the pride system to balance the competing demands of effectiveness and economy, to avoid the dual costs of under-deploying and over-deploying its outputs. To test this hypothesis, we measured people's responses regarding each of 25 socially valued traits. We observed the predicted magnitude matchings. The intensities of the pride feeling and of various motivations of pride (communicating the achievement, demanding better treatment, investing in the valued trait and pursuing new challenges) vary in proportion: (a) to one another; and (b) to the degree to which audiences value each achievement. These patterns of magnitude matching were observed both within and between the USA and India. These findings suggest that pride works cost-effectively, promoting the pursuit of achievements and facilitating the gains from others' valuations that make those achievements worth pursuing.

16.
Evol Psychol ; 19(2): 14747049211016009, 2021.
Artigo em Inglês | MEDLINE | ID: mdl-34060370

RESUMO

The synchronized co-activation of multiple responses-motivational, behavioral, and physiological-has been taken as a defining feature of emotion. Such response coherence has been observed inconsistently however, and this has led some to view emotion programs as lacking biological reality. Yet, response coherence is not always expected or desirable if an emotion program is to carry out its adaptive function. Rather, the hallmark of emotion is the capacity to orchestrate multiple mechanisms adaptively-responses will co-activate in stereotypical fashion or not depending on how the emotion orchestrator interacts with the situation. Nevertheless, might responses cohere in the general case where input variables are specified minimally? Here we focus on shame as a case study. We measure participants' responses regarding each of 27 socially devalued actions and personal characteristics. We observe internal and external coherence: The intensities of felt shame and of various motivations of shame (hiding, lying, destroying evidence, and threatening witnesses) vary in proportion (i) to one another, and (ii) to the degree to which audiences devalue the disgraced individual-the threat shame defends against. These responses cohere both within and between the United States and India. Further, alternative explanations involving the low-level variable of arousal do not seem to account for these results, suggesting that coherence is imparted by a shame system. These findings indicate that coherence can be observed at multiple levels and raise the possibility that emotion programs orchestrate responses, even in those situations where coherence is low.


Assuntos
Emoções , Vergonha , Humanos , Motivação
17.
Proc Biol Sci ; 277(1699): 3509-18, 2010 Nov 22.
Artigo em Inglês | MEDLINE | ID: mdl-20554544

RESUMO

Recent research has shown that humans, like many other animals, have a specialization for assessing fighting ability from visual cues. Because it is probable that the voice contains cues of strength and formidability that are not available visually, we predicted that selection has also equipped humans with the ability to estimate physical strength from the voice. We found that subjects accurately assessed upper-body strength in voices taken from eight samples across four distinct populations and language groups: the Tsimane of Bolivia, Andean herder-horticulturalists and United States and Romanian college students. Regardless of whether raters were told to assess height, weight, strength or fighting ability, they produced similar ratings that tracked upper-body strength independent of height and weight. Male voices were more accurately assessed than female voices, which is consistent with ethnographic data showing a greater tendency among males to engage in violent aggression. Raters extracted information about strength from the voice that was not supplied from visual cues, and were accurate with both familiar and unfamiliar languages. These results provide, to our knowledge, the first direct evidence that both men and women can accurately assess men's physical strength from the voice, and suggest that estimates of strength are used to assess fighting ability.


Assuntos
Adaptação Fisiológica/fisiologia , Percepção Auditiva/fisiologia , Adolescente , Adulto , Agressão/fisiologia , Evolução Biológica , Feminino , Força da Mão , Humanos , Masculino , Aptidão Física , Comportamento Social , Adulto Jovem
18.
Nat Hum Behav ; 4(5): 506-516, 2020 05.
Artigo em Inglês | MEDLINE | ID: mdl-32094508

RESUMO

Laws against wrongdoing may originate in justice intuitions that are part of universal human nature, according to the adaptationist theory of the origins of criminal law. This theory proposes that laws can be traced to neurocognitive mechanisms and ancestral selection pressures. According to this theory, laypeople can intuitively recreate the laws of familiar and unfamiliar cultures, even when they lack the relevant explicit knowledge. Here, to evaluate this prediction, we conduct experiments with Chinese and Sumerian laws that are millennia old; stimuli that preserve in fossil-like form the legal thinking of ancient lawmakers. We show that laypeople's justice intuitions closely match the logic and content of those archaic laws. We also show covariation across different types of justice intuitions: interpersonal devaluation of offenders, judgements of moral wrongness, mock-legislated punishments and perpetrator shame-suggesting that multiple justice intuitions may be regulated by a common social-evaluative psychology. Although alternative explanations of these findings are possible, we argue that they are consistent with the assumption that the origin of criminal law is a cognitively sophisticated human nature.


Assuntos
Direito Penal/história , Adulto , China , Crime/legislação & jurisprudência , Eutanásia/legislação & jurisprudência , Feminino , História do Século XXI , História Antiga , Humanos , Índia , Masculino , Mesopotâmia , Estados Unidos
19.
R Soc Open Sci ; 7(5): 191922, 2020 May.
Artigo em Inglês | MEDLINE | ID: mdl-32537196

RESUMO

Are pride and shame adaptations for promoting the benefits of being valued and limiting the costs of being devalued, respectively? Recent findings indicate that the intensities of anticipatory pride and shame regarding various potential acts and traits track the degree to which fellow community members value or disvalue those acts and traits. Thus, it is possible that pride and shame are engineered to activate in proportion to others' valuations. Here, we report the results of two preregistered replications of the original pride and shame reports (Sznycer et al. 2016 Proc. Natl Acad. Sci. USA 113, 2625-2630. (doi:10.1073/pnas.1514699113); Sznycer et al. 2017 Proc. Natl Acad. Sci. USA 114, 1874-1879. (doi:10.1073/pnas.1614389114)). We required the data to meet three criteria, including frequentist and Bayesian replication measures. Both replications met the three criteria. This new evidence invites a shifting of prior assumptions about pride and shame: these emotions are engineered to gain the benefits of being valued and avoid the costs of being devalued.

20.
Proc Biol Sci ; 276(1656): 575-84, 2009 Feb 07.
Artigo em Inglês | MEDLINE | ID: mdl-18945661

RESUMO

Selection in species with aggressive social interactions favours the evolution of cognitive mechanisms for assessing physical formidability (fighting ability or resource-holding potential). The ability to accurately assess formidability in conspecifics has been documented in a number of non-human species, but has not been demonstrated in humans. Here, we report tests supporting the hypothesis that the human cognitive architecture includes mechanisms that assess fighting ability-mechanisms that focus on correlates of upper-body strength. Across diverse samples of targets that included US college students, Bolivian horticulturalists and Andean pastoralists, subjects in the US were able to accurately estimate the physical strength of male targets from photos of their bodies and faces. Hierarchical linear modelling shows that subjects were extracting cues of strength that were largely independent of height, weight and age, and that corresponded most strongly to objective measures of upper-body strength-even when the face was all that was available for inspection. Estimates of women's strength were less accurate, but still significant. These studies are the first empirical demonstration that, for humans, judgements of strength and judgements of fighting ability not only track each other, but accurately track actual upper-body strength.


Assuntos
Adaptação Fisiológica/genética , Agressão/fisiologia , Face , Percepção Visual/genética , Adolescente , Adulto , Idoso , Feminino , Humanos , Masculino , Pessoa de Meia-Idade , Aptidão Física , Seleção Genética , Comportamento Social , Percepção Visual/fisiologia
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