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The rarity of Pongo fossils with precise absolute dating from the Middle Pleistocene hampers our understanding of the taxonomy and spatiotemporal distribution of Quaternary orangutans in southern China. Here, we report a newly discovered sample of 113 isolated teeth of fossil Pongo from Zhongshan Cave in the Bubing Basin, Guangxi, southern China. We describe the Pongo specimens from Zhongshan Cave and compare them metrically to other samples of fossil Pongo species (i.e., Pongo weidenreichi, Pongo devosi, Pongo duboisi, Pongo palaeosumatrensis, Pongo javensis, and Pongo sp.) and to extant orangutans (i.e., Pongo pygmaeus and Pongo abelii). The Zhongshan Pongo assemblage is dated using U-series and coupled electron spin resonance/U-series methods. Our results reasonably constrain the Zhongshan Pongo assemblage to 184 ± 16 ka, which is consistent with the biostratigraphic evidence. The Zhongshan Pongo teeth are only 6.5% larger on average than those of extant Pongo. The Zhongshan teeth are smaller overall than those of Pongo from all other cave sites in southern China, and they currently represent the smallest fossil orangutans in southern China. Based on their dental size, and the presence of a well-developed lingual pillar and lingual cingulum on the upper and lower incisors, an intermediate frequency of lingual cingulum remnants on the upper molars, and a higher frequency of moderate to heavy wrinkling on the upper and lower molars, we provisionally assign the Zhongshan fossils to P. devosi. Our results confirm earlier claims that P. weidenreichi is replaced by a smaller species in southern China, P. devosi, by the late Middle Pleistocene. The occurrence of P. devosi in Zhongshan Cave further extends its spatial and temporal distribution. The Pongo specimens from Zhongshan provide important new evidence to demonstrate that the dental morphological features of Pongo in southern China changed substantially during the late Middle Pleistocene.
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Hominidae , Pongo abelii , Diente , Animales , Pongo/anatomía & histología , Fósiles , China , Diente/anatomía & histología , Pongo pygmaeus , Hominidae/anatomía & histologíaRESUMEN
The Pongo fossil record of China extends from the Early Pleistocene to the Late Pleistocene, but to date, no late Middle Pleistocene samples of Pongo with precise absolute dating have been identified in southern China. Here, we report the recovery of 106 fossil teeth of Pongo from Ganxian Cave in the Bubing Basin, Guangxi, southern China. We dated the speleothems using Uranium-series and dated the two rhinoceros teeth using coupled electron spin resonance/Uranium-series dating methods to between 168.9 ± 2.4 ka and 362 ± 78 ka, respectively. These dates are consistent with the biostratigraphic and magnetostratigraphic age estimates. We further describe the fossil teeth from Ganxian Cave and compare them metrically to samples of fossil Pongo (i.e., Pongo weidenreichi, Pongo duboisi, Pongo palaeosumatrensis, Pongo javensis, and Pongo sp.) from the Early, Middle, and Late Pleistocene and to extant Pongo (i.e., Pongo pygmaeus and Pongo abelii) from Southeast Asia. Based on overall dental size, a high frequency of lingual cingulum remnants on the upper molars, and a low frequency of moderate to heavy wrinkling on the molars, we attribute the Ganxian fossils to P. weidenreichi. Compared with Pongo fossils from other mainland Southeast Asia sites, those from Ganxian confirm that dental size reduction of Pongo occurred principally during the Early and Middle Pleistocene. From the Middle to Late Pleistocene, all teeth except the P3 show little change in occlusal area, indicating that the size of these teeth remained relatively stable over time. The evolutionary trajectory of the Pongo dentition through time may be more complex than previously thought. More orangutan fossils with precise dating constraints are the keys to solving this issue.
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Hominidae , Pongo abelii , Uranio , Animales , Pongo , Pongo pygmaeus , China , Diente Molar , FósilesRESUMEN
Previous studies have suggested that the Lower-to-Middle Paleolithic transition was associated with the earliest Neanderthals, but recent research has established that the oldest Neanderthal fossils and the first signs of their technologies and behavior appear from MIS 11 or possibly earlier. To understand these changes, re-evaluation of the evidence is necessary to determine if this transition corresponds to a progressive evolution rather than abrupt change. Orgnac 3 is a key and appropriate site to study this research context. Located in southern France, it yields a long stratigraphic sequence testifying the evolution of technical and subsistence behaviors of pre-Neanderthal human groups during a Middle Pleistocene interglacial-glacial cycle. In this article, a new chronological framework is provided for the sequence based on results of dating methods applied to various types of geological materials. Speleothems and volcanic minerals, dated in previous studies by U-series and 40Ar/39Ar, respectively, show periods of calcitic crystallization and regional volcanic activity. Other materials, such as heated flints and herbivore teeth, are directly related to evidence of anthropogenic activities and are analyzed in the present work by trapped-charge dating methods such as thermoluminescence and electron spin resonance combined with uranium series (ESR/U-series). The new thermoluminescence and ESR/U-series dates confirm the attribution of the Orgnac 3 stratigraphic sequence to the MIS 10-MIS 8 period and are discussed in relation to paleoenvironmental data derived from bioarchaeological studies. The paleoanthropological levels, including the emergence of Levallois technology, are dated to ca. 275 ka (early MIS 8) and appear coeval to a wet and temperate period recorded locally, the Amargiers interstadial, defined in the regional palynological records. The implications of this reassessed chronology for the archaeological assemblages are discussed in the wider context of behavioral innovations from MIS 11 onward and their establishment in subsequent periods.
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Hombre de Neandertal , Diente , Animales , Arqueología , Fósiles , Francia , Humanos , TecnologíaRESUMEN
Some areas in Western Europe indicate hiatuses in human occupations, which cannot be systematically attributed to taphonomic factors and poor site preservation. The site of la Noira in the center of France records two occupation phases with a significant time gap. The older one is dated to around 700 ka (stratum a) with an Acheulean assemblage, among the earliest in Western Europe, and the upper phase of the sequence (stratum c) is dated to ca. 450 ka. Humans left the area at around 670 ka, at the beginning of the marine isotope stage (MIS) 16 glacial stage, when cold conditions became too severe. No sites between 650 and 450 ka have yet been discovered in the center region despite systematic surveys over the past three decades. The archaeological evidence indicates that populations returned to the area, at the end of MIS 12 or the beginning of the long interglacial MIS 11. Here, we use technological behaviors common to the two levels of la Noira-strata a and c to evaluate their differences. Compared to other key European sequences, this site can be used to address the evolution of the behavioral strategies in Europe between MIS 17 and 11. We formulate two hypotheses concerning the human settlement of this area: (1) local behavioral evolution over time of populations occasionally occupying the region when the climate was favorable or (2) dispersal and arrival of new populations from other areas. The results focus on (1) changes in land-use patterns with the extension of the territory used by hominins in the upper level, (2) the introduction of new core technologies, including some evidence of early Levallois debitage, and (3) more intensive shaping of bifaces and bifacial tools. Results attest that the la Noira archaeological assemblages record similar regional behavioral evolution as observed at a larger scale in Europe.
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Arqueología , Hominidae , Ocupaciones/historia , Animales , Francia , Historia Antigua , Humanos , Comportamiento del Uso de la HerramientaRESUMEN
The site of LuneryRosieres la-Terre-des-Sablons (Lunery, Cher, France) comprises early evidence of human occupation in mid-latitudes in Western Europe. It demonstrates hominin presence in the Loire River Basin during the Early Pleistocene at the transition between an interglacial stage and the beginning of the following glacial stage. Three archaeological levels sandwiched and associated with two diamicton levels deposited on the downcutting river floor indicate repeated temporary occupations. Lithic material yields evidence of simple and more complex core technologies on local Jurassic siliceous rocks and Oligocene millstone. Hominins availed of natural stone morphologies to produce flakes with limited preparation. Some cores show centripetal management and a partially prepared striking platform. The mean ESR age of 1175 ka ± 98 ka obtained on fluvial sediments overlying the archaeological levels could correspond to the transition between marine isotopic stages (MIS) 37 and 36, during the normal Cobb Mountain subchron, and in particular at the beginning of MIS 36. The Lunery site shows that hominins were capable of adapting to early glacial environmental conditions and adopting appropriate strategies for settling in mid-latitude zones. These areas cannot be considered as inhospitable at that time as Lunery lies at some distance from the forming ice cap.
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Arqueología , Sedimentos Geológicos , Humanos , Sedimentos Geológicos/análisis , Francia , Tecnología/historia , Animales , Fósiles , Hominidae , Ocupaciones/historia , Europa (Continente)RESUMEN
The Sierra de Atapuerca, northern Spain, is known from many prehistoric and palaeontological sites documenting human prehistory in Europe. Three major sites, Gran Dolina, Galería and Sima del Elefante, range in age from the oldest hominin of Western Europe dated to 1.1 to 1.3 Ma (millions of years ago) at Sima del Elefante to c.a. 0.2 Ma on the top of the Galería archaeological sequence. Recently, a chronology based on luminescence methods (Thermoluminescence [TL] and Infrared Stimulated Luminescence [IRSL]) applied to cave sediments was published for the Gran Dolina and Galería sites. The authors proposed for Galería an age of 450 ka (thousands of years ago) for the units lower GIII and GII, suggesting that the human occupation there is younger than the hominid remains of Sima de los Huesos (>530 ka) around 1 km away. In this paper, we present new results obtained by combined Electron Spin Resonance/Uranium-series (ESR/U-series) dating on 20 herbivorous teeth from different levels at the Galería site. They are in agreement with the TL results for the upper part of the stratigraphic sequence (GIV and GIIIb), in the range of between 200 and 250 ka. But for the GIIIa to GIIb levels, the TL ages become abruptly older by 200 ka while ESR ages remain relatively constant. Finally, the TL and ESR data agree in the lowest part of the section (GIIa); both fall in the range of around 350-450 ka. Our results suggest a different interpretation for the GII, GIII and GIV units of Galería and the upper part of Gran Dolina (TD10 and TD11) than obtained by TL. The ESR/U-series results are supported by a Bayesian analysis, which allows a better integration between stratigraphic information and radiometric data.
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Espectroscopía de Resonancia por Spin del Electrón/métodos , Fósiles , Mediciones Luminiscentes/métodos , Datación Radiométrica/métodos , Diente/química , Animales , Teorema de Bayes , Cronología como Asunto , Sedimentos Geológicos/química , Hominidae , Caballos , Rumiantes , España , Uranio/químicaRESUMEN
The Mauer mandible, holotype of Homo heidelbergensis, was found in 1907 in fluvial sands deposited by the Neckar River 10 km southeast of Heidelberg, Germany. The fossil is an important key to understanding early human occupation of Europe north of the Alps. Given the associated mammal fauna and the geological context, the find layer has been placed in the early Middle Pleistocene, but confirmatory chronometric evidence has hitherto been missing. Here we show that two independent techniques, the combined electron spin resonance/U-series method used with mammal teeth and infrared radiofluorescence applied to sand grains, date the type-site of Homo heidelbergensis at Mauer to 609 ± 40 ka. This result demonstrates that the mandible is the oldest hominin fossil reported to date from central and northern Europe and raises questions concerning the phyletic relationship of Homo heidelbergensis to more ancient populations documented from southern Europe and in Africa. We address the paleoanthropological significance of the Mauer jaw in light of this dating evidence.
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Fósiles , Historia Antigua , Hominidae/anatomía & histología , Datación Radiométrica/métodos , Animales , Espectroscopía de Resonancia por Spin del Electrón , Alemania , Humanos , Rayos Infrarrojos , Mandíbula/anatomía & histología , Mandíbula/diagnóstico por imagen , Radiografía , Diente/anatomía & histología , Diente/diagnóstico por imagen , UranioRESUMEN
Long considered on the margins, far from the major cultural traditions, the Sechura Desert is situated at the crossroads between the cultures of southern Ecuador and those of the northern Peruvian coast and preserves a large number of varied archaeological sites. Despite this evidence, little is known about the societies that inhabited this region during the Holocene. Exposed to natural hazards, including El Niño events, and to major climatic changes, they were able to adapt and exploit the scarce resources that this extreme environment offered them. Because of this rich history, we have been conducting archaeological research in this region since 2012 in order to clarify the dynamics of human occupation and their links with climate oscillations and environmental changes. This paper present the results of a multidisciplinary study of Huaca Grande, a mound located on Nunura Bay, 300 m from the Pacific Ocean. The nature of the human occupations at Huaca Grande was varied, and several adjustments occurred over time. The subsistence economy was based mainly on local marine resources and a continual use of terrestrial vegetal resources. However, a major change occurred in the more recent occupations, with the apparition of non-local resources (maize and cotton) indicating that Huaca Grande was connected to trade networks. The results show two main phases of occupation separated by a long abandonment (mid-5th century CE to mid-7th century CE and mid-13th century to mid-15th century CE). The occupation of the site appears to have been influenced by changes in the local climate and by extreme El Niño events. Our results highlight the great adaptability of these human groups over the span of a millennium and their capacity to react to the climatic changes and hazards that characterise this region.
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Bahías , Cambio Climático , Humanos , Perú , Océano Pacífico , EcuadorRESUMEN
After the last interglacial [Marine Isotope Stage (MIS) 5e] Europe was affected by several harsh climatic oscillations. In this context southern Italy acted, like the rest of peninsular Mediterranean Europe, as a 'glacial refugium', allowing the survival of various species, and was involved in the spread of 'cold taxa' (e.g. woolly mammoth and woolly rhino) only during the coldest phases (MIS 4 and MIS 2). Both late Mousterian and early Upper Palaeolithic sites testify to a human occupation continuity in southern Italy and especially in Apulia in this time span. Here we present a focus on three key Apulian Palaeolithic sequences (Grotta di Santa Croce, Riparo L'Oscurusciuto and Grotta del Cavallo - layers F-E) jointly spanning from the late MIS 4 to the demise of Neanderthals around 43 ka. Novel chronological, sedimentological and zooarchaeological data are discussed for the first time in the light of the palaeoenvironmental information provided by recent analyses carried out on a speleothem from Pozzo Cucù cave (Bari) and the results of the magnetic susceptibility analysis from Riparo L'Oscurusciuto. This integrated reading allows a better understanding of the role played by the Apulian region as both a refugium for late Neaderthals and a suitable habitat for the early settling of modern humans.
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The Chauvet cave (UNESCO World Heritage site, France) is located in the Ardèche Gorge, a unique physical and cultural landscape. Its setting within the gorge-overlooking a meander cutoff containing a natural arch called the Pont d'Arc-is also remarkable. Investigating possible associations between sites' physical and cultural settings, chronologies of human occupation, and access conditions has become a major theme in archeological research. The present study aims to reconstruct the landscape of the Pont d'Arc meander cutoff during the Upper Paleolithic, when humans were present in the Chauvet Cave. We used uranium-series and electron spin resonance analyses to date the formation of the Pont d'Arc natural arch in the Combe d'Arc meander cutoff, near the Chauvet Cave. Results show that the meander became totally cutoff between 108 and 138 ka (95%). Hence, the natural arch formed before the Upper Paleolithic and the first known human presence in the Chauvet Cave, dated to 37 ka cal BP. These results allowed us to reconstruct a key part of the landscape surrounding the Chauvet Cave when it was being used by Upper-Paleolithic societies.
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New Caledonia was, until recently, considered an old continental island harbouring a rich biota with outstanding Gondwanan relicts. However, deep marine sedimentation and tectonic evidence suggest complete submergence of the island during the latest Cretaceous to the Paleocene. Molecular phylogenies provide evidence for some deeply-diverging clades that may predate the Eocene and abundant post-Oligocene colonisation events. Extinction and colonization biases, as well as survival of some groups in refuges on neighbouring paleo-islands, may have obscured biogeographic trends over long time scales. Fossil data are therefore crucial for understanding the history of the New Caledonian biota, but occurrences are sparse and have received only limited attention. Here we describe five exceptional fossil assemblages that provide important new insights into New Caledonia's terrestrial paleobiota from three key time intervals: prior to the submersion of the island, following re-emergence, and prior to Pleistocene climatic shifts. These will be of major importance for elucidating changes in New Caledonia's floristic composition over time.
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Notarchirico (Southern Italy) has yielded the earliest evidence of Acheulean settlement in Italy and four older occupation levels have recently been unearthed, including one with bifaces, extending the roots of the Acheulean in Italy even further back in time. New 40Ar/39Ar on tephras and ESR dates on bleached quartz securely and accurately place these occupations between 695 and 670 ka (MIS 17), penecontemporaneous with the Moulin-Quignon and la Noira sites (France). These new data demonstrate a very rapid expansion of shared traditions over Western Europe during a period of highly variable climatic conditions, including interglacial and glacial episodes, between 670 and 650 (i.e., MIS17/MIS16 transition). The diversity of tools and activities observed in these three sites shows that Western Europe was populated by adaptable hominins during this time. These conclusions question the existence of refuge areas during intense glacial stages and raise questions concerning understudied migration pathways, such as the Sicilian route.
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The dispersal of hominin groups with an Acheulian technology and associated bifacial tools into northern latitudes is central to the debate over the timing of the oldest human occupation of Europe. New evidence resulting from the rediscovery and the dating of the historic site of Moulin Quignon demonstrates that the first Acheulian occupation north of 50°N occurred around 670-650 ka ago. The new archaeological assemblage was discovered in a sequence of fluvial sands and gravels overlying the chalk bedrock at a relative height of 40 m above the present-day maximal incision of the Somme River and dated by ESR on quartz to early MIS 16. More than 260 flint artefacts were recovered, including large flakes, cores and five bifaces. This discovery pushes back the age of the oldest Acheulian occupation of north-western Europe by more than 100 ka and bridges the gap between the archaeological records of northern France and England. It also challenges hominin dispersal models in Europe showing that hominins using bifacial technology, such as Homo heidelbergensis, were probably able to overcome cold climate conditions as early as 670-650 ka ago and reasserts the importance of the Somme valley, where Prehistory was born at the end of the 19th century.
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[This corrects the article DOI: 10.1371/journal.pone.0218591.].
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The study of geomagnetic excursions is key for understanding the behavior of the magnetic field of the Earth. In this paper, we present the geomagnetic record in a 2.29-m-long continuous core sampled in a flowstone in Liguria (Italy) and dated to the Lower Brunhes. The cored flowstone developed from Marine Isotopic Stage (MIS) 13 to MIS 7, according to 21 U-series dates. The mean growth rate is closely related to glacial and interglacial isotopic stages. Magnetic remanence was measured using u-channel and deconvolved. Four geomagnetic excursions were recorded at the same location, in a single flowstone, during interglacial MIS 11 and 13; Basura 1, 2, 3 and 4, at depths of 213 cm, 181, 160 and 92 cm, respectively. Due to the uncertainties of U-Th dating, the timing of the three events, namely Basura 1, 2 and 3 overlaps. The Basura 4 is well-dated to 417 + -7/8 ka and is clearly distinguishable from the others. It should therefore be considered as a possible excursion.
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For the past decade, debates on the earliest evidence of bifacial shaping in Western Europe have focused on several key issues, such as its origin (i.e., local or introduced), or on what should define the Acheulean culture. Whatever hypotheses are proposed for its origin, the onset and technological strategies for making Large Cutting Tools (LCTs), including biface production, are key issues and are often associated with other behavioural changes, such as increased core technology complexity. Current archaeological patterns do not support the existence of transitional industries. Rather, the scant evidence suggests that biface production associated with the management of bifacial volume was widespread around 700 ka. Among the earliest sites, the site of Notarchirico in Southern Italy stands out as one of the most significant examples. 40Ar/39Ar ages and ESR dates recently provided a revised chronology for the whole sedimentary sequence and constrained the archaeological levels between ca. 610 and 670 ka. Five archaeosurfaces (A, A1, B, D and F) yielded LCTs, including bifaces, during Marcello Piperno's excavations from 1980 to 1995. In light of this new chronological framework, which is much shorter than previously thought, we propose in this contribution a revision of the bifaces by applying the "chaine opératoire" method for the first time (analysis of reduction processes). Our goals are to assess biface production in this early Western European locality and to characterize the strategies applied at the site throughout the sequence. A corpus of 32 tools was selected from the A-A1, B, D and F archaeosurfaces. The technological analysis shows that hominins had the capacity to manage bifacial volumes, when raw material quality was adequate. Clear differences do not emerge between the different levels in terms of shaping modes or final forms. However, we demonstrate that the oldest level (level F), with the richest corpus, lacks flint and displays a higher diversity of bifaces. This ability to manage bifacial and bilateral equilibrium, as well as the diversity of the morphological results, is observed in a few penecontemporaneous sites (700-600 ka), both in the north-western and southern parts of Western Europe. These patterns suggest that hominins mastered well-controlled and diversified biface production, combining intense shaping and minimal shaping, and shared a common technological background regardless of the geographical area, and applied this technology regardless of the available raw materials. The degree of skill complexity of hominins in Western Europe between 700 and 600 ka, the current lack of evidence suggesting "gradual industries" between core-and-flake series and Acheulean techno-complexes, raise numerous questions on the origin of new behaviours in Western Europe, their mode of diffusion, and their association with Homo heidelbergensis or other Middle Pleistocene populations.
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In the last few decades, new discoveries have pushed the beginning of the biface-rich European Acheulian from 500 thousand years (ka) ago back to at least 700 ka, and possibly to 1 million years (Ma) ago. It remains, however, unclear to date if handaxes arrived in Europe as a fully developed technology or if they evolved locally from core-and-flake industries. This issue is also linked with another long-standing debate on the existence and behavioral, cognitive, and social meaning of a possibly chronological trend for increased handaxe symmetry throughout the Lower Paleolithic. The newly discovered sites can provide a link between the much older Acheulian in Africa and the Levant and the well-known assemblages from the later European Acheulian, enabling a rigorous testing of these hypotheses using modern morphometric methods. Here we use the Continuous Symmetry Measure (CSM) method to quantify handaxe symmetry at la Noira, a newly excavated site in central France, which features two archaeological levels, respectively ca. 700 ka and 500 ka old. In order to provide a context for the new data, we use a large aggregate from the well-known 500 ka old site of Boxgrove, England. We show that handaxes from the oldest layer at la Noira, although on average less symmetric than both those from the younger layers at the same site and than those from Boxgrove, are nevertheless much more symmetric than other early Acheulian specimens evaluated using the CSM method. We also correlate trends in symmetry to degree of reduction, demonstrating that raw material availability and discard patterns may affect observed symmetry values. We conclude that it is likely that, by the time the Acheulian arrived in Europe, its makers were, from a cognitive and motor-control point of view, already capable of producing the symmetric variant of this technology.
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Artrópodos , Fósiles , Animales , Arqueología , Europa (Continente) , Francia , GeografíaRESUMEN
Goda Buticha is a cave site near Dire Dawa in southeastern Ethiopia that contains an archaeological sequence sampling the late Pleistocene and Holocene of the region. The sedimentary sequence displays complex cultural, chronological and sedimentological histories that seem incongruent with one another. A first set of radiocarbon ages suggested a long sedimentological gap from the end of Marine Isotopic Stage (MIS) 3 to the mid-Holocene. Macroscopic observations suggest that the main sedimentological change does not coincide with the chronostratigraphic hiatus. The cultural sequence shows technological continuity with a late persistence of artifacts that are usually attributed to the Middle Stone Age into the younger parts of the stratigraphic sequence, yet become increasingly associated with lithic artifacts typically related to the Later Stone Age. While not a unique case, this combination of features is unusual in the Horn of Africa. In order to evaluate the possible implications of these observations, sedimentological analyses combined with optically stimulated luminescence (OSL) were conducted. The OSL data now extend the radiocarbon chronology up to 63 ± 7 ka; they also confirm the existence of the chronological gap between 24.8 ± 2.6 ka and 7.5 ± 0.3 ka. The sedimentological analyses suggest that the origin and mode of deposition were largely similar throughout the whole sequence, although the anthropic and faunal activities increased in the younger levels. Regional climatic records are used to support the sedimentological observations and interpretations. We discuss the implications of the sedimentological and dating analyses for understanding cultural processes in the region.
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Arqueología , Fósiles/diagnóstico por imagen , Sedimentos Geológicos/análisis , Datación Radiométrica/métodos , Animales , Radioisótopos de Carbono , Cuevas , Etiopía , Fósiles/historia , Historia Antigua , Humanos , Luminiscencia , Mediciones LuminiscentesRESUMEN
We present here the results of a technological and typological analysis of the Acheulian and early Middle Paleolithic assemblages from Torre in Pietra (Latium, Italy) together with comparisons with the Acheulian small tools of Castel di Guido. The assemblages were never chronometrically dated before. We have now 40Ar/39Ar dates and ESR-U-series dates, within a geomorphological framework, which support correlations to marine isotope stages. The Acheulian (previously correlated to MIS 9) is now dated to MIS 10 while the Middle Paleolithic is dated to MIS 7. Lithic analyses are preceded by taphonomic evaluations. The Levallois method of the Middle Paleolithic assemblage is an innovation characterized by the production of thin flake blanks without cortex. In contrast, the small tool blanks of the Acheulian were either pebbles or thick flakes with some cortex. They provided a relatively easy manual prehension. The choice of Levallois thin flake blanks in the Middle Paleolithic assemblage suggest that the new technology is most likely related to the emergence of hafting. Accordingly, the oldest direct evidence of hafting technology is from the site of Campitello Quarry in Tuscany (Central Italy) where birch-bark tar, found on the proximal part of two flint flakes, is dated to the end of MIS 7. Nevertheless, a peculiar feature of the Middle Paleolithic at Torre in Pietra is the continuous presence of small tool blanks on pebbles and cores and on thick flake albeit at a much lower frequency than in the older Acheulian industries. The adoption of the new technology is thus characterized by innovation combined with a degree of stability. The persistence of these habits in spite of the introduction of an innovative technique underlies the importance of cultural transmission and conformity in the behavior of Neandertals.