Root mass vertical distribution of perennial cool-season grasses grown in pure or mixed swards / Distribuição vertical da massa de raízes de gramíneas perenes de inverno cultivadas puras e em mistura

In this study we tested whether the root biomass of mixtures composed by grass species is greater than their respective monocultures. The treatments were monocultures of Arrhenatherum elatius, Festuca arundinacea, Dactylis glomerata, and a mixture of them, cultivated in a rich-soil environment. Root biomass was evaluated on a single evaluation per season at three soil depths (0-5, 5-10, and 10-20 cm). Mixed swards presented the greatest root biomass, and this was explained by a greater concentration in the topsoil layer (0-5 cm). These findings reinforce the need for permanent soil conservation practices to not jeopardize the benefits of the more abundant root biomass reported in the mixed swards.

O presente estudo testou se a biomassa radicular de misturas compostas por gramíneas é maior do que os seus respectivos monocultivos. Os tratamentos foram monocultivos de Arrhenatherum elatius, Festuca arundinacea, Dactylis glomerata e uma mistura delas, cultivadas em um solo rico em nutrientes. A biomassa da raiz foi avaliada uma vez por estação em três profundidades de solo (0-5, 5-10 e 10-20 cm). Os pastos mistos apresentaram a maior biomassa radicular e isso pode ser explicado pela maior concentração de raízes na camada superficial do solo (0-5 cm). Esses achados reforçam a necessidade de práticas permanentes de conservação do solo para não prejudicar os benefícios da biomassa radicular mais abundante encontrada nos pastos mistos.

Grazing intensity drives plant diversity but does not affect forage production in a natural grassland dominated by the tussock-forming grass Andropogon lateralis Nees.

Andropogon lateralis is a tall and highly plastic tussock-forming grass native from southern South America. It is a frequent component of Campos and Subtropical highland grasslands that often becomes dominant under lax grazing regimes. The aim of this work was to analyze the response of species diversity and forage production of a natural grassland dominated by A. lateralis to a wide range of grazing intensity. We hypothesized that species diversity and forage production would both peak at the intermediate canopy heights determined by grazing regimes of moderate intensity. A grazing experiment was conducted in a highland grassland with mesothermal humid climate at 922 masl (Atlantic Forest biome, Santa Catarina state, Brazil) that comprised 87 species from 20 families but had 50% of its standing biomass accounted by A. lateralis. Four pre-/post-grazing canopy heights-12/7, 20/12, 28/17, and 36/22 cm (measured on A. lateralis)-were arranged in a complete randomized block design with four replications, and intermittently stocked with beef heifers from October 2015 to October 2017. Andropogon lateralis cover decreased (from 75 to 50%), and species richness increased (15-25 species m-2) as canopy height decreased. Grazing intensity did not affect annual forage production (4.2 Mg DM ha-1). This natural grassland dominated by A. lateralis had a high capacity to adjust to grazing regimes of contrasting intensity, maintaining forage production stable over a wide range of canopy heights. However, to prevent losses in floristic diversity, such grassland should not be grazed at canopy heights higher than 28 cm.

Defoliation intensity and leaf area index recovery in defoliated swards: implications for forage accumulation

This study evaluated the leaf area index (LAI) recovery mechanisms and forage accumulation rates on the regrowth of different grass species subjected to different defoliation intensities. For that purpose, plots of Pennisetum clandestinum (kikuyugrass), Lolium multiflorum (annual ryegrass), and Avena strigosa (black oats) were defoliated from 20 to 80 % of their initial heights (25, 20, and 25 cm, respectively). At different increments in height, forage samples were collected to ground level and used to estimate tiller population density (TPD), leaf area per tiller (LA), and forage mass. From these data, we calculated the leaf area index (LAI), average leaf area index (aLAI), and average and instantaneous forage accumulation rate (FAR and IFAR, respectively). Data were plotted over time (days) to describe LAI recovery and forage accumulation rates. As the defoliation intensity increased, greater canopy heights were needed for pastures to achieve their maximum forage accumulation rates, which required longer regrowth intervals. The need for high tiller recruitment after defoliation, which delayed canopy LAI recovery, seemed to be one of the main cause. Thus, grazing management strategies that involve costly tiller recruitment could decrease both overall forage production and sward persistence over time. However, the plant ability to recover LAI after successive intense defoliations seems to be species-dependent and related to their phenotypic plasticity.

Defoliation intensity and leaf area index recovery in defoliated swards: implications for forage accumulation

This study evaluated the leaf area index (LAI) recovery mechanisms and forage accumulation rates on the regrowth of different grass species subjected to different defoliation intensities. For that purpose, plots of Pennisetum clandestinum (kikuyugrass), Lolium multiflorum (annual ryegrass), and Avena strigosa (black oats) were defoliated from 20 to 80 % of their initial heights (25, 20, and 25 cm, respectively). At different increments in height, forage samples were collected to ground level and used to estimate tiller population density (TPD), leaf area per tiller (LA), and forage mass. From these data, we calculated the leaf area index (LAI), average leaf area index (aLAI), and average and instantaneous forage accumulation rate (FAR and IFAR, respectively). Data were plotted over time (days) to describe LAI recovery and forage accumulation rates. As the defoliation intensity increased, greater canopy heights were needed for pastures to achieve their maximum forage accumulation rates, which required longer regrowth intervals. The need for high tiller recruitment after defoliation, which delayed canopy LAI recovery, seemed to be one of the main cause. Thus, grazing management strategies that involve costly tiller recruitment could decrease both overall forage production and sward persistence over time. However, the plant ability to recover LAI after successive intense defoliations seems to be species-dependent and related to their phenotypic plasticity.(AU)

Agronomic traits of dual-purpose wheat with different plant architectures under defoliation strategies / Características agronômicas de trigo duplo-propósito com diferentes arquiteturas de plantas sob estratégias de desfolhação

Crop-livestock system implies in the balance of appropriate animal foraging, minimizing effects on grain production on the same crop season. This study aims to assess canopy heights (20 and 30 cm) and cuts number (no cut, 1, 2 and 3 cuts) effects on wheat productivity and bromatological composition of forage, and grain yield of BRS Umbu and BRS Tarumã cultivars. Regardless cuts number and cultivars, 30 cm provided higher forage productivity (1032 and 1348 kg DM ha-1) than 20 cm (554 and 909 kg DM ha-1), for BRS Umbu and Tarumã, respectively, in average crop seasons. For both cultivars and crop seasons, mean values of 26% crude protein, 55% neutral detergent fiber and 26% acid detergent fiber were obtained. At 20 cm no difference in grain yield was observed as a function of cuts number with 3780 kg ha-1 for BRS Umbu, in average crop seasons and 4306 kg ha-1 in 2014, for BRS Tarumã. However, at 30 cm, the grain yield was penalized at least in 30% as a result only third cut. These results indicate that wheat as dual-purpose can be employed in southern Brazil in grazing systems in forage shortage times and then produce grains without yield penalties as a function to the management used.

A integração lavoura-pecuária implica no equilíbrio do forrageamento adequado dos animais, minimizando os impactos nas áreas agrícolas graníferas. Os objetivos deste trabalho foram verificar, em trigo, os efeitos das alturas pré-corte (20 e 30 cm) e do número de cortes (sem corte, 1, 2 e 3 cortes) sobre a produtividade e composição bromatológica da forragem, e a produtividade de grãos dos cultivares BRS Umbu e BRS Tarumã. Independente do número de cortes, a altura de 30 cm proporcionou maior produtividade de forragem (1032 e 1348 kg MS ha-1) do que 20 cm (554 e 909 kg MS ha-1), para os cultivares BRS Umbu e BRS Tarumã, respectivamente, na média das safras. Para ambos cultivares e safras, obteve-se valores médios de 26% de proteína bruta, 55% de fibra em detergente neutro e 26% de fibra em detergente ácido. A 20 cm não foram encontradas diferenças na produtividade de grãos em função do número de cortes, com 3780 kg ha-1 para BRS Umbu na média das safras e 4306 kg ha-1 em 2014 para BRS Tarumã. Porém, para ambos os cultivares a 30 cm, o rendimento de grãos foi prejudicado em 30% no terceiro corte. Estes resultados indicam que o trigo duplo-propósito pode ser empregado no sul do Brasil em sistemas de pastejo em épocas de escassez de forragem e ainda produzir grãos sem penalidades, de acordo com o manejo empregado.

Milk fatty acid composition of unsupplemented dairy cows grazing on a tropical pasture

This study aimed to evaluate the milk fatty acid (FA) composition of unsupplemented dairy cows grazing on elephantgrass (Pennisetum purpureum Schum. cv. Pioneiro) pastures under three grazing management strategies: pre-grazing height of 120 cm with defoliation intensities of 50 and 70% of the initial height (treatments 120/50 and 120/70) and pre-grazing height of 90 cm with defoliation intensity of 70% (treatment 90/70). Treatments were repeated three times in a complete randomized block design. Tester animals were six multiparous Holsteins cows grouped into pairs according to days in milk and milk yield and allocated to blocks. Individual milk samples were collected after the first and second days of grazing (two-day occupation period) and analyzed for FA composition. Milk fat from 120/50 and 90/70 treatments resulted in higher concentrations of C4:0, C6:0, C17:0 anteiso, rumenic acid (RA), vacenic acid (VA), trans-12 C18:1, trans-13/14 C18:1, cis-11 C18:1, cis-12 C18:1, cis-9, trans-12 C18:2, trans-11, cis-15 C18:2, and total polyunsaturated fatty acids. Additionally, both SCD14 (cis-9 14:1/14:0 + cis-9 14:1) and SCDRA (RA/VA + RA) desaturase indices were reduced in milk fat from cows subjected to 120/50 and 90/70 treatments. On average, the milk fat levels of RA and VA observed in our study were greater than those reported in most studies with tropical grass-based diets and similar to levels usually found in cows grazing on temperate pastures. Overall, our results showed that milk with a desirable fatty acid composition is produced by cows grazing on a tropical grass (elephantgrass cv. Pioneiro), with the combination of a pre-grazing height of 120 cm and a defoliation intensity of 50% of the initial height, being a practical management strategy to achieve this goal.(AU)

Experimental evidence that the perennial grass persistence pathway is linked to plant growth strategy.

Grass species can be classified into different functional types based on their growth strategies, and contrasting persistence strategies are observed in different grass species. Excluding seedling recruitments, changes in populations of grasses are basically a trade-off between natality and mortality of tillers. We hypothesised that the persistence pathway of perennial grasses is linked to their growth strategy, regardless whether they are growing as monoculture or as a mixture. Species with contrasting growth strategies (Arrhenatherum elatius L., Dactylis glomerata L., and Festuca arundinacea Schreb.) were cultivated as monocultures and as a mixture and their tiller natality and mortality were evaluated for two years after swards establishment. All pastures maintained their population size during the experimental period, although decreases in tiller densities occurred during the warmer season. Arrhenatherum elatius had the highest tiller mortality and natality whereas the F. arundinacea had the lowest ones. Arrhenatherum elatius had many tillers appearing in all seasons but their tillers were short-lived. Conversely, F. arundinacea and D. glomerata developed numerous tillers during autumn and winter and their tillers survived, on average, almost six and three times longer than those of A. elatius, respectively. There were no differences in tillering dynamics among populations grown in monocultures or in the mixture. Regardless of whether they were cultivated in monocultures or as a mixture, the persistence pathway of perennial grasses is linked with their growth strategies with exploitative species presenting a high tiller turnover throughout the year whereas the persistence of more conservative species is based on a high tiller survival.

Limestone and phosphorus application and forage production in natural pastures with sodseeding of cool-season species / Aplicação de calcário e fósforo e a produção de forragem em pastagem natural com introdução de espécies de estação fria

Natural pastures are important ecosystems that both contribute to biodiversity conservation and provide an important source of income, especially for cattle ranchers. While these pastures yield fairly low productivity, they can be improved by increasing soil fertility and introducing species with high productive potentials. In this sense, the purpose of this study was to evaluate the effects of applying limestone and phosphorus, as well as introducing cool-season species with a mixture of species dominated by Schizachyrium tenerum Nees, into a natural pasture in the Catarinense Plateau. The experiment was conducted from January 2010 to December 2013; the treatments consisted of superficial distribution of limestone in proportions of 3.6, 7.2, 11.0, and 14.4 t ha-1, as well as distribution of phosphorus in the form of triple superphosphate in proportions of 35, 70, 105, and 140 kg of P2O5 ha-1. In addition, cool-season species were overseeded. The experiment consisted of a randomized block design with subdivided plots and three replications. Limestone was applied to the main parcel, whereas phosphorus was applied to the subplots. There was no interaction between the levels of limestone and phosphorus. The application of 11.0 t ha-1 of limestone yielded the highest forage production, with 3,932.2 kg of dry matter (DM) ha-1 during the second year. Red clover was the species that best reacted to the additions, with levels of 7.2 and 11.0 t ha-1 over the 4 years. In addition, phosphorus provoked a positive response throughout the experiment. The highest forage production was observed during the second year, with an addition of 140 kg P2O5 ha-1 (4,419.4 kg DM ha-1).

As pastagens naturais representam um importante ecossistema, com contribuições tanto para manutenção da biodiversidade quanto para geração de renda aos produtores, principalmente na pecuária de corte. No entanto, os níveis de produtividade destas pastagens são relativamente baixos, e podem ser melhorados com incrementos em fertilidade do solo e introdução de espécies de elevado potencial produtivo. Assim, o objetivo deste estudo foi avaliar os efeitos da aplicação de níveis de calcário e fósforo, bem como da introdução de espécies de inverno em uma pastagem natural com predomínio de Schizachyrium tenerum Nees no Planalto Catarinense. O experimento foi conduzido de janeiro de 2010 a dezembro de 2013, e os tratamentos consistiram na distribuição superficial de calcário nas doses de 3,6; 7,2; 11,0 e 14,4 t ha-1; distribuição de fósforo na forma de superfosfato triplo nas doses de 35; 70; 105 e 140 kg de P2O5 ha-1; e a sobressemeadura de espécies de estação fria. O delineamento experimental foi em blocos ao acaso com parcelas subdivididas e três repetições. Os níveis de calcário foram distribuídos nas parcelas e os de fósforo nas subparcelas. Não houve interação entre os níveis de calcário e fosforo. A aplicação de 11,0 t ha-1 de calcário propocionou a maior produção de forragem, de 3.932,2 kg MS ha-1, no 2º ano.

Limestone and phosphorus application and forage production in natural pastures with sodseeding of cool-season species / Aplicação de calcário e fósforo e a produção de forragem em pastagem natural com introdução de espécies de estação fria

Natural pastures are important ecosystems that both contribute to biodiversity conservation and provide an important source of income, especially for cattle ranchers. While these pastures yield fairly low productivity, they can be improved by increasing soil fertility and introducing species with high productive potentials. In this sense, the purpose of this study was to evaluate the effects of applying limestone and phosphorus, as well as introducing cool-season species with a mixture of species dominated by Schizachyrium tenerum Nees, into a natural pasture in the Catarinense Plateau. The experiment was conducted from January 2010 to December 2013; the treatments consisted of superficial distribution of limestone in proportions of 3.6, 7.2, 11.0, and 14.4 t ha-1, as well as distribution of phosphorus in the form of triple superphosphate in proportions of 35, 70, 105, and 140 kg of P2O5 ha-1. In addition, cool-season species were overseeded. The experiment consisted of a randomized block design with subdivided plots and three replications. Limestone was applied to the main parcel, whereas phosphorus was applied to the subplots. There was no interaction between the levels of limestone and phosphorus. The application of 11.0 t ha-1 of limestone yielded the highest forage production, with 3,932.2 kg of dry matter (DM) ha-1 during the second year. Red clover was the species that best reacted to the additions, with levels of 7.2 and 11.0 t ha-1 over the 4 years. In addition, phosphorus provoked a positive response throughout the experiment. The highest forage production was observed during the second year, with an addition of 140 kg P2O5 ha-1 (4,419.4 kg DM ha-1).(AU)

As pastagens naturais representam um importante ecossistema, com contribuições tanto para manutenção da biodiversidade quanto para geração de renda aos produtores, principalmente na pecuária de corte. No entanto, os níveis de produtividade destas pastagens são relativamente baixos, e podem ser melhorados com incrementos em fertilidade do solo e introdução de espécies de elevado potencial produtivo. Assim, o objetivo deste estudo foi avaliar os efeitos da aplicação de níveis de calcário e fósforo, bem como da introdução de espécies de inverno em uma pastagem natural com predomínio de Schizachyrium tenerum Nees no Planalto Catarinense. O experimento foi conduzido de janeiro de 2010 a dezembro de 2013, e os tratamentos consistiram na distribuição superficial de calcário nas doses de 3,6; 7,2; 11,0 e 14,4 t ha-1; distribuição de fósforo na forma de superfosfato triplo nas doses de 35; 70; 105 e 140 kg de P2O5 ha-1; e a sobressemeadura de espécies de estação fria. O delineamento experimental foi em blocos ao acaso com parcelas subdivididas e três repetições. Os níveis de calcário foram distribuídos nas parcelas e os de fósforo nas subparcelas. Não houve interação entre os níveis de calcário e fosforo. A aplicação de 11,0 t ha-1 de calcário propocionou a maior produção de forragem, de 3.932,2 kg MS ha-1, no 2º ano. (AU)

White grubs (Cyclocephala flavipennis) damaging perennial winter pastures in the South Region of Brazil / Ocorrência do coró-pequeno (Cyclocephala flavipennis) danificando pastagens perenes de inverno na região Sul do Brasil

Cyclocephala flavipennis Arrow, 1914 (Coleoptera: Melolonthidae), popularly known as white grub, is an insect widely disseminated in the South Region of Brazil. Despite the frequent occurrence of this white grub in areas cultivated with winter cereals, it is generally not considered a pest, since it feeds on straw and dead plant material. However, in 2015 and 2016, this insect has been identified as the damage-causing agent in the perennial winter pastures in Lages, Santa Catarina, Brazil. Damage was observed in patches and caused reduction in root depth and mass, resulting in loss of vigor, accelerated senescence in aerial parts, and death of tillers in Festuca arundinacea Schreb. and Dactylis glomerata L. plants.

Cyclocephala flavipennis Arrow, 1914 (Coleoptera: Melolonthidae), conhecido popularmente como coró-pequeno, é um inseto amplamente disseminado na região sul do Brasil. Embora seja encontrado com frequência em áreas cultivadas com cereais de inverno não é considerado inseto praga, por alimentar-se preferencialmente de palha e restos de plantas. Entretanto, nos anos de 2015 e 2016 foi identificado como causador de danos em pastagens perenes de inverno em Lages, Santa Catarina, Brasil. Os danos foram observados em reboleiras e promoveram redução na massa e profundidade de raízes, resultando em perda de vigor, senescência acelerada da parte aérea e morte de perfilhos de plantas de Festuca arundinacea Schreb. e Dacytilis glomerata L.

White grubs (Cyclocephala flavipennis) damaging perennial winter pastures in the South Region of Brazil / Ocorrência do coró-pequeno (Cyclocephala flavipennis) danificando pastagens perenes de inverno na região Sul do Brasil

Cyclocephala flavipennis Arrow, 1914 (Coleoptera: Melolonthidae), popularly known as white grub, is an insect widely disseminated in the South Region of Brazil. Despite the frequent occurrence of this white grub in areas cultivated with winter cereals, it is generally not considered a pest, since it feeds on straw and dead plant material. However, in 2015 and 2016, this insect has been identified as the damage-causing agent in the perennial winter pastures in Lages, Santa Catarina, Brazil. Damage was observed in patches and caused reduction in root depth and mass, resulting in loss of vigor, accelerated senescence in aerial parts, and death of tillers in Festuca arundinacea Schreb. and Dactylis glomerata L. plants.(AU)

Cyclocephala flavipennis Arrow, 1914 (Coleoptera: Melolonthidae), conhecido popularmente como coró-pequeno, é um inseto amplamente disseminado na região sul do Brasil. Embora seja encontrado com frequência em áreas cultivadas com cereais de inverno não é considerado inseto praga, por alimentar-se preferencialmente de palha e restos de plantas. Entretanto, nos anos de 2015 e 2016 foi identificado como causador de danos em pastagens perenes de inverno em Lages, Santa Catarina, Brasil. Os danos foram observados em reboleiras e promoveram redução na massa e profundidade de raízes, resultando em perda de vigor, senescência acelerada da parte aérea e morte de perfilhos de plantas de Festuca arundinacea Schreb. e Dacytilis glomerata L.(AU)

White grubs (Cyclocephala flavipennis) damaging perennial winter pastures in the South Region of Brazil / Ocorrência do coró-pequeno (Cyclocephala flavipennis) danificando pastagens perenes de inverno na região Sul do Brasil

ABSTRACT: Cyclocephala flavipennis Arrow, 1914 (Coleoptera: Melolonthidae), popularly known as white grub, is an insect widely disseminated in the South Region of Brazil. Despite the frequent occurrence of this white grub in areas cultivated with winter cereals, it is generally not considered a pest, since it feeds on straw and dead plant material. However, in 2015 and 2016, this insect has been identified as the damage-causing agent in the perennial winter pastures in Lages, Santa Catarina, Brazil. Damage was observed in patches and caused reduction in root depth and mass, resulting in loss of vigor, accelerated senescence in aerial parts, and death of tillers in Festuca arundinacea Schreb. and Dactylis glomerata L. plants.

RESUMO: Cyclocephala flavipennis Arrow, 1914 (Coleoptera: Melolonthidae), conhecido popularmente como coró-pequeno, é um inseto amplamente disseminado na região sul do Brasil. Embora seja encontrado com frequência em áreas cultivadas com cereais de inverno não é considerado inseto praga, por alimentar-se preferencialmente de palha e restos de plantas. Entretanto, nos anos de 2015 e 2016 foi identificado como causador de danos em pastagens perenes de inverno em Lages, Santa Catarina, Brasil. Os danos foram observados em reboleiras e promoveram redução na massa e profundidade de raízes, resultando em perda de vigor, senescência acelerada da parte aérea e morte de perfilhos de plantas de Festuca arundinacea Schreb. e Dacytilis glomerata L.

Forage accumulation during the stocking period on pastures subjected to intermittent stocking / Acúmulo de forragem durante o período de ocupação dos animais em pastos sob lotação intermitente

The aim of this study was to evaluate the forage accumulation during the stocking period on Kikuyu grass pastures (Pennisetum clandestinum Hochst. Ex. Chiov) subjected to intermittent stocking. Treatments consisted of four defoliation levels (removal of 40, 50, 60, and 70% relative to the initial height) starting from the same pre-grazing height (20 cm). The following variables were evaluated: leaf and stem elongation rates (evaluated during four times across the period of occupation), leaf area index, and forage accumulation rates. Additionally, ten tillers/strip were evaluated for the same variables into exclusion cages. The linear, quadratic and cubic effects of severity of defoliation were evaluated by orthogonal polynomials contrasts with 5% of significance level. There was a linear and inverse relationship between the proportion of removed leaf area index with leaf elongation and herbage accumulation rate. The leaf elongation rate in individual tillers and the forage accumulation rates of tillers on excluded areas from grazing were greater than those under grazing. During the stocking period on pastures submitted to intermittent grazing, forage accumulation was observed and it was linear and negatively correlated with the proportion of leaf area removed.

O objetivo deste trabalho foi avaliar o acúmulo de forragem durante o período de ocupação pelos animais em pastos de capim-quicuiu (Pennisetum clandestinum Hochst. Ex. Chiov.) manejados sob lotação intermitente. Os tratamentos consistiram de quatro proporções de desfolhação (remoção de 40, 50, 60 e 70% da altura inicial) partindo de uma mesma altura em pré-pastejo (20 cm). Foram avaliadas: taxas de alongamento foliar e colmos e senescência de folhas, índice de área foliar e taxa de acúmulo de forragem. Adicionalmente, dez perfilhos por faixa foram avaliados para as mesmas variáveis dentro de gaiolas de exclusão. Os efeitos linear, quadrático e cúbico da severidade de desfolhação foram avaliados por contrastes de polinômios ortogonais com 5% de nível de significância. Houve relação linear e inversa entre a porcentagem do índice de área foliar removido e o alongamento de lâminas foliares e o acúmulo de forragem. As taxas de alongamento de folhas em perfilhos bem como o acúmulo de forragem nas áreas excluídas do pastejo foram maiores em relação aos pastos submetidos a desfolhação. Durante o período de rebaixamento de pastos manejados sob lotação intermitente ocorre acúmulo de forragem e o mesmo é linear e inversamente associado à proporção da área foliar removida.

Forage accumulation during the stocking period on pastures subjected to intermittent stocking / Acúmulo de forragem durante o período de ocupação dos animais em pastos sob lotação intermitente

The aim of this study was to evaluate the forage accumulation during the stocking period on Kikuyu grass pastures (Pennisetum clandestinum Hochst. Ex. Chiov) subjected to intermittent stocking. Treatments consisted of four defoliation levels (removal of 40, 50, 60, and 70% relative to the initial height) starting from the same pre-grazing height (20 cm). The following variables were evaluated: leaf and stem elongation rates (evaluated during four times across the period of occupation), leaf area index, and forage accumulation rates. Additionally, ten tillers/strip were evaluated for the same variables into exclusion cages. The linear, quadratic and cubic effects of severity of defoliation were evaluated by orthogonal polynomials contrasts with 5% of significance level. There was a linear and inverse relationship between the proportion of removed leaf area index with leaf elongation and herbage accumulation rate. The leaf elongation rate in individual tillers and the forage accumulation rates of tillers on excluded areas from grazing were greater than those under grazing. During the stocking period on pastures submitted to intermittent grazing, forage accumulation was observed and it was linear and negatively correlated with the proportion of leaf area removed.(AU)

O objetivo deste trabalho foi avaliar o acúmulo de forragem durante o período de ocupação pelos animais em pastos de capim-quicuiu (Pennisetum clandestinum Hochst. Ex. Chiov.) manejados sob lotação intermitente. Os tratamentos consistiram de quatro proporções de desfolhação (remoção de 40, 50, 60 e 70% da altura inicial) partindo de uma mesma altura em pré-pastejo (20 cm). Foram avaliadas: taxas de alongamento foliar e colmos e senescência de folhas, índice de área foliar e taxa de acúmulo de forragem. Adicionalmente, dez perfilhos por faixa foram avaliados para as mesmas variáveis dentro de gaiolas de exclusão. Os efeitos linear, quadrático e cúbico da severidade de desfolhação foram avaliados por contrastes de polinômios ortogonais com 5% de nível de significância. Houve relação linear e inversa entre a porcentagem do índice de área foliar removido e o alongamento de lâminas foliares e o acúmulo de forragem. As taxas de alongamento de folhas em perfilhos bem como o acúmulo de forragem nas áreas excluídas do pastejo foram maiores em relação aos pastos submetidos a desfolhação. Durante o período de rebaixamento de pastos manejados sob lotação intermitente ocorre acúmulo de forragem e o mesmo é linear e inversamente associado à proporção da área foliar removida.(AU)

ACÚMULO DE FORRAGEM DURANTE O PERÍODO DE OCUPAÇÃO DOS ANIMAIS EM PASTOS SOB LOTAÇÃO INTERMITENTE

Abstract The aim of this study was to evaluate the forage accumulation during the stocking period on Kikuyu grass pastures (Pennisetum clandestinum Hochst. Ex. Chiov) subjected to intermittent stocking. Treatments consisted of four defoliation levels (removal of 40, 50, 60, and 70% relative to the initial height) starting from the same pre-grazing height (20 cm). The following variables were evaluated: leaf and stem elongation rates (evaluated during four times across the period of occupation), leaf area index, and forage accumulation rates. Additionally, ten tillers/strip were evaluated for the same variables into exclusion cages. The linear, quadratic and cubic effects of severity of defoliation were evaluated by orthogonal polynomials contrasts with 5% of significance level. There was a linear and inverse relationship between the proportion of removed leaf area index with leaf elongation and herbage accumulation rate. The leaf elongation rate in individual tillers and the forage accumulation rates of tillers on excluded areas from grazing were greater than those under grazing. During the stocking period on pastures submitted to intermittent grazing, forage accumulation was observed and it was linear and negatively correlated with the proportion of leaf area removed.

Resumo O objetivo deste trabalho foi avaliar o acúmulo de forragem durante o período de ocupação pelos animais em pastos de capim-quicuiu (Pennisetum clandestinum Hochst. Ex. Chiov.) manejados sob lotação intermitente. Os tratamentos consistiram de quatro proporções de desfolhação (remoção de 40, 50, 60 e 70% da altura inicial) partindo de uma mesma altura em pré-pastejo (20 cm). Foram avaliadas: taxas de alongamento foliar e colmos e senescência de folhas, índice de área foliar e taxa de acúmulo de forragem. Adicionalmente, dez perfilhos por faixa foram avaliados para as mesmas variáveis dentro de gaiolas de exclusão. Os efeitos linear, quadrático e cúbico da severidade de desfolhação foram avaliados por contrastes de polinômios ortogonais com 5% de nível de significância. Houve relação linear e inversa entre a porcentagem do índice de área foliar removido e o alongamento de lâminas foliares e o acúmulo de forragem. As taxas de alongamento de folhas em perfilhos bem como o acúmulo de forragem nas áreas excluídas do pastejo foram maiores em relação aos pastos submetidos a desfolhação. Durante o período de rebaixamento de pastos manejados sob lotação intermitente ocorre acúmulo de forragem e o mesmo é linear e inversamente associado à proporção da área foliar removida.

Forage production in a natural grassland with limestone and phosphorus dosages / Produção de forragem em pastagem natural submetida a aplicações de calcário e fósforo

The objective of this study was to assess forage production in a natural grassland by applying different doses of limestone and phosphorus, and the influence of these applications on soil profile. The treatments consisted of the application of limestone doses of 0.0, 7.2, and 14.4 t ha-1 and phosphorus doses of 0,35, 70, and 140 kg of P2O5 ha-1 on the soil surface. The experiment was conducted in a randomized complete block design with split-plots and three replications. The limestone doses were distributed in the main plot and applied only at the beginning of the experiment, whereas the phosphorus doses were distributed in the subplots with subsequent annual maintenance applications. Was used a ruler (cm) for monitoring the growth of pastures and, when grasses reached an average height of 20 cm, forage samples were collected in each subplot and botanical and morphological components were separated and dried until a constant weight was achieved. In the third and fourth assessment years, soil samples were collected in the 0-5, 5-10, 10-15, and 15-20-cm layers to evaluate the influence of treatments on the chemical properties of the soil. The variables analyzed were related to the production of several components, including fodder, native grasses, leguminous plants, unwanted plants, and dead material, in addition to soil properties such as pH, exchangeable aluminum concentration, and base saturation. There was no interaction between the limestone and phosphorus treatments. The effect of limestone doses on forage production was assessed in the third year, and, in the following year, the intermediate dose of limestone (7.2 t ha-1) produced the highest yield (2,316.1 kg of dry matter [DM] of forage ha-1).

O objetivo deste trabalho foi avaliar a produção de forragem em pastagem natural submetida a doses superficiais de calcário e fósforo e a influência dessas aplicações ao longo do perfil do solo. Os tratamentos consistiram da aplicação de doses superficiais de calcário equivalentes a 0,0; 7,2 e 14,4 t.ha-1 e de fósforo equivalentes a 0; 35; 70 e 140 kg de P2O5 ha-1. O experimento foi conduzido de acordo com um delineamento em blocos completos casualizados com parcelas subdivididas e três repetições. As doses de calcário foram distribuídas na parcela principal e aplicadas unicamente no início do experimento enquanto que as doses de fósforo foram distribuídas nas subparcelas com posteriores aplicações anuais de manutenção. O acompanhamento do crescimento dos pastos foi realizado por meio de uma régua graduada em centímetros e, quando os pastos atingiam altura média de 20 cm, eram coletadas amostras de forragem de cada subparcela e posteriormente os componentes botânicos e morfológicos eram separados e secos até atingirem peso constante. No 3 e 4 ano de avaliação foram coletadas amostras de solo nas camadas de 0-5, 5-10, 10-15 e 15-20 cm de profundidade para verificar a influência dos tratamentos nos atributos químicos do solo. As variáveis analisadas envolveram desde a produção de forragem, de gramíneas e leguminosas nativas, de plantas indesejáveis e de material morto e atributos do solo como pH, concentração de alumínio trocável e saturação por bases. Não houve interação entre os tratamentos de calcário e fósforo. O efeito das doses de calcário sobre a produção de pastagem natural ocorreu a partir do 3º ano e, no ano seguinte, a dose intermediária do corretivo (7,2 t.ha-1) foi responsável pelas maiores produtividades (2.316,1 kg de MS de forragem.ha-1). As crescentes doses de fósforo aumentaram a produção de forragem a partir do 2° ano, atingindo produções de até 2.232 kg MS.ha-1 no 4° ano.

Forage production in a natural grassland with limestone and phosphorus dosages / Produção de forragem em pastagem natural submetida a aplicações de calcário e fósforo

The objective of this study was to assess forage production in a natural grassland by applying different doses of limestone and phosphorus, and the influence of these applications on soil profile. The treatments consisted of the application of limestone doses of 0.0, 7.2, and 14.4 t ha-1 and phosphorus doses of 0,35, 70, and 140 kg of P2O5 ha-1 on the soil surface. The experiment was conducted in a randomized complete block design with split-plots and three replications. The limestone doses were distributed in the main plot and applied only at the beginning of the experiment, whereas the phosphorus doses were distributed in the subplots with subsequent annual maintenance applications. Was used a ruler (cm) for monitoring the growth of pastures and, when grasses reached an average height of 20 cm, forage samples were collected in each subplot and botanical and morphological components were separated and dried until a constant weight was achieved. In the third and fourth assessment years, soil samples were collected in the 0-5, 5-10, 10-15, and 15-20-cm layers to evaluate the influence of treatments on the chemical properties of the soil. The variables analyzed were related to the production of several components, including fodder, native grasses, leguminous plants, unwanted plants, and dead material, in addition to soil properties such as pH, exchangeable aluminum concentration, and base saturation. There was no interaction between the limestone and phosphorus treatments. The effect of limestone doses on forage production was assessed in the third year, and, in the following year, the intermediate dose of limestone (7.2 t ha-1) produced the highest yield (2,316.1 kg of dry matter [DM] of forage ha-1). (AU)

O objetivo deste trabalho foi avaliar a produção de forragem em pastagem natural submetida a doses superficiais de calcário e fósforo e a influência dessas aplicações ao longo do perfil do solo. Os tratamentos consistiram da aplicação de doses superficiais de calcário equivalentes a 0,0; 7,2 e 14,4 t.ha-1 e de fósforo equivalentes a 0; 35; 70 e 140 kg de P2O5 ha-1. O experimento foi conduzido de acordo com um delineamento em blocos completos casualizados com parcelas subdivididas e três repetições. As doses de calcário foram distribuídas na parcela principal e aplicadas unicamente no início do experimento enquanto que as doses de fósforo foram distribuídas nas subparcelas com posteriores aplicações anuais de manutenção. O acompanhamento do crescimento dos pastos foi realizado por meio de uma régua graduada em centímetros e, quando os pastos atingiam altura média de 20 cm, eram coletadas amostras de forragem de cada subparcela e posteriormente os componentes botânicos e morfológicos eram separados e secos até atingirem peso constante. No 3 e 4 ano de avaliação foram coletadas amostras de solo nas camadas de 0-5, 5-10, 10-15 e 15-20 cm de profundidade para verificar a influência dos tratamentos nos atributos químicos do solo. As variáveis analisadas envolveram desde a produção de forragem, de gramíneas e leguminosas nativas, de plantas indesejáveis e de material morto e atributos do solo como pH, concentração de alumínio trocável e saturação por bases. Não houve interação entre os tratamentos de calcário e fósforo. O efeito das doses de calcário sobre a produção de pastagem natural ocorreu a partir do 3º ano e, no ano seguinte, a dose intermediária do corretivo (7,2 t.ha-1) foi responsável pelas maiores produtividades (2.316,1 kg de MS de forragem.ha-1). As crescentes doses de fósforo aumentaram a produção de forragem a partir do 2° ano, atingindo produções de até 2.232 kg MS.ha-1 no 4° ano.(AU)

A grazing height target to minimize tiller stem elongation rate in annual ryegrass swards / Altura limite de pastejo para minimizar o alongamento de colmos em pastos de azevém anual

ABSTRACT: Italian ryegrass (Lolium multiflorum Lam) is one of the most cultivated temperate annual forage crop around the world. Despite that, there is little information about the process of stem elongation during its vegetative growth. This information would be useful for grazing management purposes. Thus, the aim of this study was to assess the herbage accumulation and stem elongation rates in annual ryegrass pastures submitted to intermittent stocking. The experimental design used was a completely randomized 2x2 factorial with three replications. The treatments were defined by the combination of two pre-grazing heights (15 and 25cm) associated with two post-grazing heights (4 and 8cm). The herbage accumulation rates were higher in pastures with 25cm pre-grazing associated with post-grazing heights of 8cm. However, leaf production rate was only affected by post-grazing height, with higher values being recorded for stubbles heights of 8cm, regardless of pre-grazing height. Stem elongation rates in annual ryegrass tillers increased rapidly when pastures reached around 18cm, a condition where there was a relatively low competition for light, suggesting that; 1) pre-grazing heights of 18 to 20cm in annual ryegrass could be used as a upper limit for grazing management purposes whether the objective is to minimize stem contribution in accumulated herbage and 2) It seems that stem elongation is as inevitable process (independently of competition for light) and that this could be related to the necessary strength of stems, pseudostems and sheaths to support larger and heavier leaves.

RESUMO: O azevém anual (Lolium multuflorum Lam) é uma das plantas forrageiras de clima temperado mais cultivadas no mundo. Apesar disso, há poucas informações sobre o processo de alongamento de colmos para essa espécie durante seu crescimento vegetativo. Tal informação seria útil para o manejo mais adequado desta espécie. Assim, o objetivo do presente trabalho foi avaliar o acúmulo de forragem e as taxas de alongamento de colmos em pastos de azevém anual submetidos à lotação intermitente. O delineamento experimental foi completamente casualizado em arranjo factorial 2x2, com três repetições. Os tratamentos consistiram na combinação de duas alturas em pré-pastejo (15 e 25cm), associadas com duas alturas de resíduo (4 e 8cm). A maior taxa de acúmulo de forragem foi observada nos pastos manejadas com 25cm, associadas com alturas de resíduo de 8cm. Entretanto, a taxa de acúmulo de folhas foi similar para pastos manejados com 8cm de resíduo, independente da altura em pré-pastejo (15 ou 25cm). A taxa de alongamento de colmos em perfilhos de azevém anual aumentaram rapidamente quando os pastos atingiram uma altura de 18cm, condição onde havia uma relativa baixa competição por luz, sugerindo que: 1) alturas em pré-pastejo entre 18 e 20cm em pastos de azevém anual poderia ser usada como um limite máximo de altura quando o objetivo for minimizar a contribuição de colmos na forragem acumulada; e 2) parece que o alongamento de colmos é um processo inevitável (independente de competição por luz) e isso poderia estar relacionado com a força necessária de colmos, pseudocolmos e bainhas para suportar o peso de folhas cada vez maiores e mais pesadas.

A grazing height target to minimize tiller stem elongation rate in annual ryegrass swards / Altura limite de pastejo para minimizar o alongamento de colmos em pastos de azevém anual

Italian ryegrass (Lolium multiflorum Lam) is one of the most cultivated temperate annual forage crop around the world. Despite that, there is little information about the process of stem elongation during its vegetative growth. This information would be useful for grazing management purposes. Thus, the aim of this study was to assess the herbage accumulation and stem elongation rates in annual ryegrass pastures submitted to intermittent stocking. The experimental design used was a completely randomized 2x2 factorial with three replications. The treatments were defined by the combination of two pre-grazing heights (15 and 25cm) associated with two post-grazing heights (4 and 8cm). The herbage accumulation rates were higher in pastures with 25cm pre-grazing associated with post-grazing heights of 8cm. However, leaf production rate was only affected by post-grazing height, with higher values being recorded for stubbles heights of 8cm, regardless of pre-grazing height. Stem elongation rates in annual ryegrass tillers increased rapidly when pastures reached around 18cm, a condition where there was a relatively low competition for light, suggesting that; 1) pre-grazing heights of 18 to 20cm in annual ryegrass could be used as a upper limit for grazing management purposes whether the objective is to minimize stem contribution in accumulated herbage and 2) It seems that stem elongation(AU)

O azevém anual (Lolium multuflorum Lam) é uma das plantas forrageiras de clima temperado mais cultivadas no mundo. Apesar disso, há poucas informações sobre o processo de alongamento de colmos para essa espécie durante seu crescimento vegetativo. Tal informação seria útil para o manejo mais adequado desta espécie. Assim, o objetivo do presente trabalho foi avaliar o acúmulo de forragem e as taxas de alongamento de colmos em pastos de azevém anual submetidos à lotação intermitente. O delineamento experimental foi completamente casualizado em arranjo factorial 2x2, com três repetições. Os tratamentos consistiram na combinação de duas alturas em pré-pastejo (15 e 25cm), associadas com duas alturas de resíduo (4 e 8cm). A maior taxa de acúmulo de forragem foi observada nos pastos manejadas com 25cm, associadas com alturas de resíduo de 8cm. Entretanto, a taxa de acúmulo de folhas foi similar para pastos manejados com 8cm de resíduo, independente da altura em pré-pastejo (15 ou 25cm). A taxa de alongamento de colmos em perfilhos de azevém anual aumentaram rapidamente quando os pastos atingiram uma altura de 18cm, condição onde havia uma relativa baixa competição por luz, sugerindo que: 1) alturas em pré-pastejo entre 18 e 20cm em pastos de azevém anual poderia ser usada como um limite máximo de altura quando o objetivo for minimizar a contribuição de colmos na forragem acumulada; e (AU)

Levels of defoliation and regrowth dynamics in elephant grass swards / Níveis de desfolhação e a dinâmica da rebrotação em pastos de capim-elefante

The morphogenetic and structural characteristics of forage plants and the processes of growth and senescence in tissues may vary according to the grazing management strategies used. This research aimed to assess the process of regrowth in elephant grass swards Cv. 'Pioneiro' submitted to two pre-grazing heights (90 and 120cm) combined with two levels of defoliation (50 and 70%). A reduction of the leaf elongation rate (LER) was observed during the regrowth stages, in addition to a reduction in the rate of leaf appearance (LAR) and aerial tiller population density (ATPD). In contrast, an increase was observed in leaf senescence (LSR) and stem and pseudo stem elongation rates (SER). The severe defoliation (70%) combined with a pre-grazing height of 120cm compromised the regrowth of elephant grass cv. 'Pioneiro', which must be managed with a maximum height of 90cm and maximum level of defoliation of 50% of the initial height.

As características morfogênicas e estruturais das plantas forrageiras, bem como os processos de crescimento e senescência de tecidos podem variar de acordo com as estratégias de manejo utilizadas. O objetivo deste trabalho foi avaliar o processo de rebrotação em pastos de capim-elefante, cultivar 'Pioneiro', submetidos a duas alturas em pré-pastejo (90 e 120cm) combinadas com duas severidades de desfolhação (50 e 70% da altura inicial). Ao longo das fases de rebrotação, ocorreu uma diminuição das taxas de alongamento foliar (TAF), aparecimento de folhas (TAP) e da densidade populacional de perfilhos aéreos (DPPA), aumento das taxas de senescência foliares (TSF) e alongamento de colmo mais pseudocolmo (TACP). A severidade de desfolha de 70% aliada à altura em pré-pastejo de 120cm comprometeu o processo de rebrotação do capim-elefante cv. 'Pioneiro', devendo este ser manejado com altura máxima de 90cm e com severidade de desfolha não excedendo 50% da altura inicial.