Clavigeritae (Coleoptera, Staphylinidae, Pselaphinae) of the Arabian Peninsula with the description of a new species of Corynotopsis Jeannel, 1951 from Oman.

Corynotopsisomanicussp. nov. of the myrmecophilous supertribe Clavigeritae is described from Oman. The type series was collected at night and the ant host species remains unknown. Corynotopsisscotti Jeannel, 1951 is newly recorded for Yemen, and Commatocerusconcinnus Besuchet & Cuccodoro, 2011 for Oman. Lasius Fabricius, 1804 and Lepisiota Santschi, 1926 are, for the first time, determined as hosts of the latter species. The problematic taxonomic status of Corynotopsisscotti is discussed. A distribution map of all Clavigeritae known from the Arabian Peninsula is provided.

Percussiopalpus gen. n. from Turkey, a free-living relative of cave-dwelling Thaumastocephalus, and revised placement of Thaumastocephalini (Coleoptera: Staphylinidae: Pselaphinae).

A new genus and species, Percussiopalpus inusitatus Hlavác & Jaloszynski gen. n. et sp. n. of the tribe Thaumastocephalini is described and illustrated, based on specimens collected in the Asian part of Turkey (Balikesir Province). The discovery of a free-living pselaphine species morphologically closely resembling the obligate cavernicolous Thaumastocephalus Poggi et al. prompted a detailed morphological study of both genera, which results in transferring Thaumastocephalini from Batrisitae to Euplectitae.

Oberprielerius, a new genus for Anchonidium uniforme Voss, 1974, a microphthalmic species of Rhythirrinini (Coleoptera: Curculionidae: Cyclominae) from South Africa.

A new genus, Oberprielerius gen. n., from the Republic of South Africa, is described for Anchonidium uniforme Voss, 1974 in the tribe Rhythirrinini Lacordaire, 1863 of the subfamily Cyclominae Schoenherr, 1826.

Pselaphinae of the Cameroon Volcanic Line. Part 1. New Pselaphinae (Coleoptera, Staphylinidae) from the island São Tomé.

A new genus of Pselaphinae, Reichentomica gen. nov., and three new species, Reichentomica tasani sp. nov., Afroplectus turneri sp. nov., and Parasimus saotomicus sp. nov., are described for the first time from the island São Tomé. All three species were collected at the same locality close to Jardim Botânico do Bom Successo (00°16'31''N, 06°36'14''E) using flight intercept trap in the tropical forest at the elevation of 1324 m.

New insights into the subterranean genus Speoplanes Müller, 1911 (Coleoptera, Leiodidae, Cholevinae) from the Dinarides.

In the present study morphological and molecular data from specimens of the troglobitic genus Speoplanes have been examined. The two subspecies Speoplanes giganteus giganteus Müller, 1911 and Speoplanes giganteus biocovensis Müller, 1934 are changed in rank from subspecies to species level and discrete identification characters for the separation of the two species are provided. Additionally, all known localities of the species (including new records) are listed and a distribution map of the genus is provided.

Endogean and Cavernicolous Coleoptera of the Balkans. XXIII. A new species of Paramaurops J. Müller (Coleoptera: Staphylinidae: Pselaphinae) from Greece.

We describe and illustrate a new species of pselaphine beetles, Paramaurops zagoricus sp. n., from the Zagori region of Epirus, Greece. A list summarising the distribution of Amauropini known from Greece is also provided.

Physico-Chemical and Sensory Profiles of Enriched Linz Biscuits.

The aim of the present study was to determine the physico-chemical properties (dry matter content, combustion heat, electrical properties, total protein, ash, fat and crude fibre contents, selected amino acids, and trace elements), antioxidant content, and sensory profile of Linz biscuits. They were enriched by the addition of powdered carrot, nettle leaves and elderberry fruit, which is 3% of the product. For comparison of results, a control variant without the addition of these components was also prepared. The enriched biscuits showed slightly higher total ash and crude fibre contents in comparison to the control samples. Results for the antioxidant activity and total polyphenol, flavonoid, and phenolic acid contents of the enriched biscuits were higher in all observed parameters than in the control sample with the best results obtained for Linz biscuits enriched with elderberry and nettle powder. In enriched biscuits, higher contents of iron, zinc, and manganese were measured, especially in biscuits with nettle. Linz biscuits with nettle had higher combustion heat values than control samples; the other two sample types had lower values. We found that the resistance, capacitance, and relative permittivity of the enriched biscuits decreased with frequency according to the power regression function. On the contrary, the conductivity increased with an increasing frequency. Electrical properties were mainly influenced by the water content but also by added components.

New synonyms of Euconnus/ (Tetramelus/) kraussi/ Reitter, 1880 (Coleoptera, Staphylinidae, Scydmaeninae).

The subgenus Tetramelus Motschulsky, 1869 with its currently 109 palaearctic species, is the largest Palaearctic subgenus of the largest world scydmaenine genus Euconnus Thomson, 1859. The nomenclatural validity of the subgenus-name has been recently supported by a detailed morphological study (Jaloszynski 2015). Euconnus (Tetramelus) kraussi was described by Edmund Reitter (1881a), with the type locality indicated as Caucasus, Martkopi. This was the second species of the subgenus Tetramelus known to occur in the entire Caucasus region, after Euconnus (Tetramelus) reitteri Saulcy, 1878, the latter with unknown type locality. The former was treated as a junior synonym of the latter by Croissandeau (1898), and as a subspecies by Karaman (1973). A modern redescription and illustration of the aedeagus was given by Vít Hlavác (1998), together with a key to all species of Tetramelus known at that time from the Caucasus. Castellini (2006: 102) placed Euconnus (Tetramelus) kraussi as a junior synonym of Euconnus (Tetramelus) reitteri. This taxonomic act was based just on the similarity of aedeagi of these two taxa illustrated by Vít Hlavác (1998). No type material was studied. Euconnus (Tetramelus) reitteri is smaller, only 1.25-1.30 mm long, against 1.40-1.60 mm for Euconnus (Tetramelus) kraussi. To combine this with differences on the aedeagus and with the allopatric distribution, we regard these two forms as two different species. Later, two new species were described, Euconnus (Tetramelus) marinae Franz, 1979, the type locality "Kachetien nächst Shuamta" and Euconnus (Tetramelus) nachuzrischwilii Franz, 1986, the type locality "nördlich von Anamuri in etwa 1000 m". Both were included in the key of Vít Hlavác (1998). The type material of both these species was found well-preserved in the Natural History Museum in Vienna (NMW), and thanks to Dr. Harald Schillhammer it was available to the senior author for study. A detailed study of this material proved that both species were conspecific with Euconnus (Tetramelus) kraussi, and their synonymy is formally established here. We are also providing new records for this species, which is so far endemic to eastern Georgia, well delimited by the Meskheti-Likhi range. The material treated in this study is deposited in the following collections: MNHN - Muséum National d'Histoire Naturelle, Paris, France; NMW - Naturhistorisches Museum Wien, Vienna, Austria; PCAP - private collection of Andreas Pütz, Eisenhüttenstadt, Germany; PCHM - private collection of Heinrich Meybohm, Großhansdorf, Germany; PCMS - private collection of Michael Schülke, Berlin, Germany; PCPH - private collection of Peter Hlavác, Prague, Czech Republic. The distributional map was created using Encarta software.

In the twilight zone-The head morphology of Bergrothia saulcyi (Pselaphinae, Staphylinidae, Coleoptera), a beetle with adaptations to endogean life but living in leaf litter.

The pselaphine Bergrothia saulcyi shows features seemingly linked with life in deep soil layers, such as greatly reduced and non-functional compound eyes, a sensorium of long tactile setae, long appendages, and flightlessness. However, the tiny beetles occur in forest leaf litter, together with a community of beetles with wings and well-developed eyes. We hypothesize that B. saulcyi moves into deep soil under dry conditions, and returns to upper layers when humidity increases again. Despite the evolutionary cost of a reduced dispersal capacity, this life strategy may be more efficient and less hazardous than moving to different habitats using flight and the visual sense in an environment periodically drying out. We also discuss cephalic features with potential phylogenetic relevance. Plesiomorphies of B. saulcyi include the presence of anterior tentorial arms, well-developed labral retractors, and a full set of extrinsic maxillary and premental muscles. Apomorphic cephalic features support clades Protopselaphinae + Pselaphinae, and Pselaphinae. A conspicuous derived condition, the clypeo-ocular carina, is a possible synapomorphy of Batrisitae and genera assigned to Goniaceritae. A complex triple set of cephalic glands found in B. saulcyi is similar to a complex identified in the strict myrmecophile Claviger testaceus (Clavigeritae). It is conceivable that glands linked with food uptake in free-living pselaphines were genetically re-programmed in ancestors of inquilines, to enable them to appease the host ants. We suggest that behavioral studies are necessary to understand the poorly known life habits of B. saulcyi. Additional information is required to explain why a species with irreversibly reduced visual sense and other adaptations typical of endogean or cave-dwelling beetles was only collected from the upper leaf litter layer.

The Pselaphinae (Coleoptera: Staphylinidae) of New Caledonia and Loyalty Islands. IV. Revision of the genus Baraxina Raffray, 1896.

The endemic New Caledonian Brachyglutini genus Baraxina is revised. Seventeen species are recognized, of which sixteen are newly described: B. francoisi Raffray, 1886; B. amieuana sp. n., B. grimbacheri sp. n., B. kanakorum sp. n., B. rutai sp. n., B. aoupinica sp. n., B. asymmetrica sp. n., B. burwelli sp. n., B. dzumacana sp. n., B. jenisi sp. n., B. koghisiana sp. n., B. lescheni sp. n., B. monteithi sp. n., B. parakoghisiana sp. n., B. pecki sp. n., B. poroana sp. n., and B. wanati sp. n.

New species of Clavigeritae (Coleoptera: Staphylinidae: Pselaphinae) from Malawi and South Africa.

Three new species of Clavigeritae from subsaharan Africa, Zuluclavodes mulanjensis sp. nov., Zuluclavodes minor sp. nov. from southern Malawi and Articerodes jaloszynskii sp. nov. from South Africa, KwaZulu-Natal are described. Diagnosis is provided for both genera, and a key to species is given for the genus Zuluclavodes Hlavác, 2007. Ischyroceros mirus Reichensperger, 1915 is firstly recorded from Malawi, the illustration of its aedeagus is given. All described species are illustrated and their photos are presented.

A second species of Syrraphesina Raffray found adjacent to Wallace's Line(Coleoptera, Staphylinidae, Pselaphinae).

The monospecific genus Syrraphesina Raffray, 1903 contains a single species distributed in eastern New Guinea. In this paper a second species from Lombok Island, S. agostii Yin Hlavác, sp. nov., is described, figured, and compared to S. pliciventris Raffray, 1903. A revised diagnosis of Syrraphesina is provided. The distribution of Clavigerini through the islands of the world is briefly discussed.

A new, peculiar genus of Cossoninae (Coleoptera, Curculionidae) from Oman with description of a new species, larva and notes on biology.

A new genus and species of the subfamily Cossoninae, Omanocossonus sabulosus gen. et sp. nov. is described from Oman. All specimens including larvae were found on sand dunes on the seashore in roots of Calotropis procera (Aiton) W.T. Aiton. The crucial features, including male and female terminalia, are illustrated, and the taxonomic position of the new genus within Cossoninae is briefly discussed. The generic status of Lindbergius Roudier, 1957 is resurrected. The mature larva of the new species is described, larval morphology is discussed and the current state of knowledge about immature stages of Cossoninae is summarized.

Endogean and Cavernicolous Coleoptera of the Balkans. XX. Notes on the genus Protamaurops J. Müller, 1944 (Coleoptera: Staphylinidae: Pselaphinae) with the description of a new species from Bulgaria.

The genus Protamaurops J. Müller, 1944 and three species: P. macedonicus J. Müller, 1934, P. macrophthalmus J. Müller, 1944 and P. winneguthi Székessy, 1943 are reviewed and redescribed. One new species from Bulgaria, Protamaurops bulgaricus sp. n. is described. Protamaurops montenegrinus (Székessy, 1943) is transferred to the genus Paramaurops Jeannel, 1948 and Protamaurops macrophtalmus beieri (J. Müller, 1944) is synonymized to the nominal species Protamaurops macrophtalmus macrophtalmus (J. Müller, 1944).

Validation of Taiwanophodes minor Hlavác, 2003 (Coleoptera: Staphylinidae: Pselaphinae).

In my 2003 paper (Hlavác 2003) dealing with a revision of the Pselaphodes complex of genera, I neglected to explicitly designate a depository of the holotype of Taiwanophodes minor Hlavác sp. n. Under ICZN (1999): Article 16.4.2 this omission invalided its description and the species name. I herein rectify this problem.

Description of new species of Pseudamaurops Jeannel, 1948 (Coleoptera: Staphylinidae: Pselaphinae) from Greece and North Macedonia.

Two new species of Pseudamaurops Jeannel, 1948 are described, P. hristovskii sp. n. from North Macedonia and P. tymficus sp. n. from Greece. Redescriptions of P. albanicus (Apfelbeck, 1907) and P. muellerianus (Ravasini, 1923) are provided, and an identification key for the genus is provided.

The new species of the genus Atanygnathus Jacobson, 1909 (Coleoptera: Styphylinidae: Staphylininae) from Réunion Island.

New species Atanygnathus poussereaui Kocian Hlavác, sp. n. from Réunion is described and illustrated. Key to all species of Atanygnathus (Coleoptera, Staphylinidae, Staphylininae, Tanygnathinina) of Réunion is presented.

A new cavernicolous species of the genus Bergrothia Reitter, 1884 (Coleoptera, Staphylinidae, Pselaphinae) from Georgia.

During a biospeleological survey of caves of Imereti region (Georgia), a new species of the beetle genus Bergrothia Reitter, B. barbakadzei sp. nov. was discovered in three caves: Datvis (Bear), Prometheus and Melouri caves (Sataplia-Tskaltubo karst massif, Tskaltubo district). The new species is described, illustrated and differentiated from other morphologically similar species of the genus-B. mingrelica (Reitter). Bergrothia barbakadzei sp. nov. is the first cavernicolous species of the genus Bergrothia. A key to the species of this genus is provided.

Endogean and cavernicolous Coleoptera of the Balkans. XVIII. Strong radiation in caves of the Central Dinarides: seven new species of Thaumastocephalus Poggi et al., 2001 (Staphylinidae: Pselaphinae).

Seven new species of the cavernicolous and anophthalmous genus Thaumastocephalus Poggi, Nonveiller, Colla, Pavicevic T. Rada, 2001 are described: T. bilandzijae sp. n., T. kirini sp. n., T. marsici sp. n., T. rujnicensis sp. n., T. slavkoi sp. n. and T. troglavi sp. n. from Croatia and T. dahnae sp. n. from Bosnia and Herzegovina. Aedeagi of all species are illustrated. A key to all species is provided. The records of all specimens of the genus treated here are given, and their distributions are discussed and shown on maps. The distribution of all genera of cavernicolous Pselaphinae in the Dinarides is discussed.

Review of the Stomodes tolutarius species group (Coleoptera: Curculionidae: Entiminae: Otiorhynchini) from the Balkan Peninsula with description of two new species from Greece.

Species of the tolutarius species group of the genus Stomodes Schoenherr, 1826 are revised. A redescription of the genus is provided and all available male and female genitalia are illustrated. Two new species, S. benedikti sp. n. and S. dodocunevi sp. n. from Greece, are described. A lectotype is designated for S. amorei Desbrochers des Loges, 1904. The redescriptions of the following species are provided: S. marocanus Hoffmann, 1956; S. muelleri Lona, 1922; S. leonhardi Wagner, 1912; S. letzneri Reitter, 1889 and S. tolutarius Schoenherr, 1826. S. muelleri is for the first time recorded for Macedonia, Bosnia and Hercegovina and Greece; S. tolutarius for Macedonia, Bulgaria and Turkey. An identification key to species of the tolutarius species group is given.