On the taxonomic status of the tenebrionoid genera Falsopedilus Pic, 1924 and Hintonosia Báguena Corella, 1948 (Coleoptera: Mycteridae) with descriptions of new species.

We re-evaluate the taxonomic placement of the genus Hintonosia Báguena Corella, 1948, hitherto placed in Aderidae Csíki, 1909. The assessment of key morphological characters based on the type specimen of the type species of this genus, Hintonosia bismacrocephalus Báguena Corella, 1948 (= Hintonosia macrocephalus Pic, 1935, homonym), demonstrated that it does not possess the synapomorphies of the Aderidae. Instead, its clear position in Mycteridae Blanchard, 1845 is uncovered, leading us to the transfer of Hintonosia to the subfamily Eurypinae Thomson, 1860 of Mycteridae. Moreover, Hintonosia was found congeneric with the mycterid genus Falsopedilus Pic, 1924 and the new synonymy is therefore proposed. Additionally, two African species of Falsopedilus are described, a key to, and a checklist of Falsopedilus species are presented.

Rediscovery of Tomosomus Motschulsky, 1855 (Coleoptera: Anthicidae) with redescription of the genus, a new genus-rank synonym, and a key to species.

Tomosomus Motschulsky, 1855, an obscure incertae sedis genus of Anthicidae erroneously considered monotypic, was found to be congeneric with the tomoderine genus Holcopyge Champion, 1890. A new genus-rank synonym (Tomosomus Motschulsky = Holcopyge Champion syn. nov.) and two new combinations are proposed for the species hitherto attributed to Holcopyge. Tomosomus subtestaceus Motschulsky, 1855, the type of its genus, is redescribed from rediscovered type material and recently collected specimens. An annotated checklist, key to species and new records of Tomosomus are provided.

Fourth contribution to the Macratriinae LeConte, 1862 of the Papuan Region and Wallacea (Coleoptera: Anthicidae) with descriptions of sixteen new species.

Descriptions of the following 16 new species of Macratriinae LeConte, 1862 are presented based on the material from various scientific institutions: Macratria anus, M. bimaculosa, M. bonggo, M. koiari, M. korowai, M. kundratai, M. longitarsis, M. perkovskyi, M. ripicola, M. roosilehti, M. saccharum, M. sentani, M. spatulatocalcarata, M. susuami, M. tigrina, M. tongkonan spp. nov. New distribution records of eight Macratria Newman, 1838 species are provided.

New species and records of Macratriinae LeConte, 1862 (Coleoptera: Anthicidae) from the Americas with five new descriptions.

Descriptions of the following five new species of Macratriinae LeConte, 1862 are presented based on the material mainly from the Natural History Museum London: Macratria bugle sp. nov. (Panama), M. conlei sp. nov. (Ecuador), M. florsavichi sp. nov. (Panama), M. guaymi sp. nov. (Panama) and M. tiriyo sp. nov. (Guyana / Suriname). New distribution records for the Central American Macratria obsoleta Champion, 1890 are also provided.

First record of the family Diplommatinidae Gray, 1847 (Gastropoda: Architaenioglossa) from Ecuador with description of a new Adelopoma Doering, 1885.

Diplommatinidae Gray, 1847, a family of minute cyclophoroid gastropods, is recorded from Ecuador for the first time. Adelopoma gracile Gree, sp. nov. is described and illustrated from River Pastaza valley, central Ecuador. Differential diagnosis is provided for similar congeners. An annotated checklist of and a preliminary key to the Adelopoma Doering, 1885 species are provided.

Redefinition of Physciolagria Pic, 1930 (Coleoptera: Mycteridae) with a new family placement, genus-rank synonymy, new description, and a key.

Physciolagria Pic, 1930, a monotypic genus erroneously attributed to Tenebrionidae incertae sedis was found to be congeneric with the eurypine genus Lacconotopedilus Pic, 1935 (Mycteridae Blanchard, 1845). The taxonomic position and comparative morphology of Physciolagria are reassessed and discussed. A new genus-rank synonym (Physciolagria Pic, 1930 = Lacconotopedilus Pic, 1935 syn. nov.), one species-rank synonym (Physciolagria liturata Pic, 1930 = Lacconotopedilus elongatus Pic, 1936 syn. nov.) and two new combinations are proposed. Physciolagria smithi sp. nov. from Senegal is described and P. liturata and P. singularicornis are redescribed and illustrated. An annotated checklist, key to species and new records are provided.

The first fossil Coleoptera record from the Volyn Region, Ukraine, with description of a new Glesoconomorphus (Coleoptera, Mycteridae) in syninclusion with Winterschmidtiidae (Acari) and a key to species.

Glesoconomorphusekaterinae sp. nov. (Coleoptera, Mycteridae), representing the first ever fossil species of Coleoptera from the Volyn Region of Ukraine and the first mycterid from late Eocene Rovno amber, is described and illustrated. A key to species of the fossil mycterid genus Glesoconomorphus Alekseev, Pollock & Bukejs, 2019 is presented. The systematic position of Glesoconomorphus within Eurypinae J. Thomson, 1860 is briefly discussed. The oldest finding of phoretic Winterschmidtiidae Oudemans, 1923 mites, found on the type specimen of the new beetle species, is reported.

Redefinition of Steriphodon Abeille de Perrin, 1895 (Coleoptera: Anthicidae), with a new species description, checklist, and key to species.

Steriphodon Abeille de Perrin, 1895, a small genus of Eurygeniinae (Anthicidae) is reviewed. Critical morphology of the genus is reassessed and discussed. An annotated checklist, key to species and new records are provided. Steriphodon ottomerkli sp. nov. from the Balochistan Province, Pakistan is described and illustrated.

The easternmost record of Macratriinae LeConte, 1862 (Coleoptera: Anthicidae), with a new species from Fiji and a genus-rank synonymy in Macratriini LeConte, 1862.

The easternmost record of Macratria Newman, 1838 from Fiji is presented, and M. fijiana sp. nov. is described and illustrated. Biogeographical patterns and diversity of Pacific Macratriinae are briefly discussed. Additionally, a new genus rank synonymy in Macratriinae is proposed: Thambospasta Werner, 1974 syn. nov. of Salimuzzamania Abdullah, 1968. New combination is made for Salimuzzamania howdeni (Werner, 1974) comb. nov. (from Thambospasta).

Review of Aderidae Csiki, 1909 (Insecta: Coleoptera) of Cabo Verde.

The Aderidae (Coleoptera: Tenebrionoidea) of Cabo Verde (or Cape Verde in English transcription) Archipelago in the Central Atlantic Ocean is revised based on an examination of types and additional material. Four species are confirmed for the archipelago, three of the genus Cobososia Collado et Alonso-Zarazaga, 1996 and one of the genus Aderus Stephens, 1829. We also propose to transfer Anthicus reductus Wollaston, 1867 (Anthicidae: Anthicinae: Anthicini) to the genus Cobososia (Aderidae), as Cobososia reducta (Wollaston, 1867) comb. nov. and demonstrate, that Cobososia angulithorax (Desbrochers des Loges, 1881) is conspecific with Cobososia reducta (Wollaston, 1867) syn. nov. Finally, we present new records and an illustrated identification key to the ant-like leaf beetles of Cabo Verde.

Addendum to Zootaxa 4683 (4): Popillia biguttata (Wiedemann in Wiedemann Germar, 1821) (Coleoptera: Scarabaeidae: Rutelinae) established on New Guinea? With an illustrated key to Sundaland Popillia Dejean species.

In our recent publication (Telnov Zorn 2019) we overlooked and did not include Popillia migliaccioi Sabatinelli, 1996, described from "Java", in our checklist and key of Sundaland Popillia species. However, the examination of the holotype of this species deposited in the Muséum d'Histoire Naturelle, Geneva, Switzerland (Figs. 1-8) revealed that Popillia migliaccioi is in fact an Afrotropical species, which was already described under the name Popillia maynei Ohaus, 1914. It is obvious that the specimen studied and described by Sabatinelli (1996) is mislabelled and is not from Java (Fig. 8). Unfortunately, the type material of P. maynei, which is housed in the Museum für Naturkunde, Berlin, Germany, is on loan indefinitely and was not available for our study (Bernd Jäger, personal communication). However, the original description of P. maynei and the aedeagus as originally figured by Ohaus (1914) (Fig. 9) match habitus and aedeagus of the P. migliaccioi holotype perfectly (Figs. 4-6). Habitus photographs of an additional comparative specimen from Togo (Kloto, III.2015, Don P. Léonard, deposited in the Royal Belgian Institute of Natural Sciences, Brussels, Belgium) are provided for comparison in figs. 10 and 11.

Climatic oscillations in Quaternary have shaped the co-evolutionary patterns between the Norway spruce and its host-associated herbivore.

During the Last Glacial Maximum in the Northern Hemisphere, expanding ice sheets forced a large number of plants, including trees, to retreat from their primary distribution areas. Many host-associated herbivores migrated along with their host plants. Long-lasting geographic isolation between glacial refugia could have been led to the allopatric speciation in separated populations. Here, we have studied whether the migration history of the Norway spruce Picea abies in Quaternary has affected its host-associated herbivorous beetle-Monochamus sartor. By using microsatellite markers accompanied by the geometric morphometrics analysis of wing venation, we have revealed the clear geographic structure of M. sartor in Eurasia, encompassing two main clusters: southern (Alpine-Carpathian) and eastern (including northeastern Europe and Asia), which reflects the northern and southern ecotypes of its host. The two beetles' lineages probably diverged during the Pleniglacial (57,000-15,000 BC) when their host tree species was undergoing significant range fragmentation and experienced secondary contact during post-glacial recolonization of spruce in the Holocene. A secondary contact of divergent lineages of M. sartor has resulted in the formation of the hybrid zone in northeastern Europe. Our findings suggest that the climatic oscillations during the Pleistocene have driven an insect-plant co-evolutionary process, and have contributed to the formation of the unique biodiversity of Europe.

Description of a new fossil Statira Lepeletier et Audinet-Serville, 1828 (Coleoptera: Tenebrionidae: Lagriinae) from Baltic amber of the Sambian Peninsula.

Statira baltica sp. nov. is described from Eocene Baltic amber found in the Baltic Sea coast, Yantarny settlement, Kaliningrad Region, Russia. This is the second described fossil member of the genus Statira Lepeletier et Audinet-Serville, 1828 and the tribe Lagriini Latreille, 1825, and the first from outside the New World.

Popillia biguttata (Wiedemann in Wiedemann amp; Germar, 1821) (Coleoptera: Scarabaeidae: Rutelinae) established on New Guinea? With an illustrated key to Sundaland Popillia Dejean species.

Of the megadiverse Old-World genus Popillia Dejean, 1821, only seven species are so far recorded from the Sunda Islands (see checklist). The ruteline scarab beetle Popillia biguttata (Wiedemann in Wiedemann Germar, 1821) was originally described from, and was until now known only from Java (Greater Sunda Islands). In 2018, specimens of this species were found in northern New Guinea, far outside its native range, in mixed autochthonous and invasive vegetation.

Gonialaenini, a new tribe of Lagriinae (Coleoptera: Tenebrionidae) from Eocene Baltic Amber.

Based on well preserved specimen, Gonialaena groehni gen. et sp. nov. (Coleoptera: Tenebrionidae) is described from Eocene Baltic amber and assigned to Gonialaenini trib. nov. within the subfamily Lagriinae due to the combination of the following characters: labrum elongate; antennal sensilla simple; procoxal cavities closed at least externally; narrow membranes present between abdominal ventrites 3-5; mesocoxal cavities partly closed by mesepimeron, trochantine present; ovipositor of primitive lagrioid type-digitate fourth coxite lobe with gonostyles apically. Gonialaena gen. nov. combines characters of various Lagriinae tribes: humeral angles rounded, formed by basally widened epipleura as in Goniaderini, Pycnocerini and Lupropini; narrow acute intercoxal process of abdominal ventrite 1 as in Belopini, Cossyphini and Lagriini; absence (or strong reduction) of metathoracic wings as in most Adeliini and some Lupropini; large, reniform eyes as in Goniaderini, Pycnocerini, Lagriini and many Adeliini; present intersegmental membranes between abdominal ventrites 3-5 as in all lagriine tribes except for Belopini and Cossyphini; subcylindrical terminal maxillary palpomere as in Belopini and Cossyphini. Thus, the position of Gonialaenini trib. nov. within Lagriinae is not fully understood, but combination of certain features (humeral angles rounded but distinct and vertical border present at elytral base, inconspicuous elytral striae, large reniform eyes, narrow intersegmental membranes) points on morphological similarity with Goniaderini, especially some Anaedus Blanchard, 1845 with indistinct elytral striae and related genera.

New species of Ischalia Pascoe, 1860 (Coleoptera: Ischaliidae), with a key to aposematically coloured species from continental Asia, and new notes on their mimicry.

Three new species of false fire-coloured beetles, namely Ischalia (s. str.) ancora, I. (s. str.) holzschuhi, and I. (s. str.) laosensis spp. n., are described from Laos and Vietnam. Ischalia (s. str.) martensi Paulus, 1971 is redescribed from the type and additional material. Male genitalia are for the first time illustrated in I. (Eupleurida) sichuanensis Young, 2008. A key to aposematically coloured species from the Asian continent is presented for the first time. Their mimicry association is further investigated and a new case involving Anthicidae is reported.

An annotated checklist of the Anthicidae and pediline Pyrochroidae (Insecta: Coleoptera) of Iran, with thirteen new country records.

The fauna of Iranian Anthicidae and pediline Pyrochroidae (Coleoptera) is summarized in this paper. A total of 132 species from 19 genera and four subfamilies (Anthicinae, Notoxinae, Steropinae, and Tomoderinae) of Anthicidae and two species of Pedilinae are listed. Three new synonyms are proposed: Anthicus armatus Truqui, 1855 = A. pseudoarmatus Telnov, 2008 syn. nov., Notoxus hirtus LaFerté-Sénectère, 1849 = N. caucasicus Pic, 1900 syn. nov., Stricticomus rufithorax (LaFerté-Sénectère, 1849) = Anthicus transcaspicus var. subnotata Pic, 1910 syn. nov. One new combination is made: Nitorus unifasciatus (Desbrochers des Loges, 1875) comb. nov. (from Pseudoleptaleus Pic, 1900). Thirteen species are reported for the fauna of Iran for the first time: Anthicus armatus Truqui, 1855, A. episcopalis Pic, 1903, A. quadrispilus Marseul, 1879, Cyclodinus reitteri (Pic, 1892), Leptaleus klugii klugii LaFerté-Sénectère, 1849, Stricticomus araxicola (Reitter, 1889), S. arcuaticeps (Pic, 1900), S. herzi (Pic, 1905), Microhoria aphaenops (Pic, 1902), Tenuicomus finalis Telnov, 2003, Notoxus hirtus LaFerté-Sénectère, 1849, N. simulans simulans Heberdey, 1935, and Tomoderus scydmaenoides Reitter, 1878.

Multiphoton Ionization of One-Electron Relativistic Diatomic Quasimolecules in Strong Laser Fields.

We perform a theoretical and computational study of relativistic one-electron homonuclear diatomic quasimolecules subject to strong electromagnetic fields linearly polarized along the molecular axis. Several quasimolecules with the nuclear charges 1-92 and appropriately scaled internuclear distances and field parameters are used in the calculations. The time-dependent Dirac equation is solved with the help of the generalized pseudospectral method in prolate spheroidal coordinates. We have found that employing this coordinate system makes it possible to avoid emergence of spurious states, which usually show up when solving the Dirac equation numerically. For lower carrier frequencies, interaction with the driving field is described within the dipole approximation. Relativistic effects in the multiphoton ionization probabilities are investigated with respect to the internuclear distance in the quasimolecule. For higher frequencies, the interaction with the field is described beyond the dipole approximation. Nondipole effects in the ionization probability are discussed.

New data on the cockroaches previously attributed to the genus Duchailluia Rehn, 1933 (Dictyoptera: Blattidae: Blattinae).

The new species of the genus Duchailluia Rehn, 1933, D. ivindo sp. nov., is described from Gabon. The genus Distylopyga Bey-Bienko, 1965 is restored from synonymy with Duchailluia. Distylopyga beccalonii sp. nov. from Thailand is described. Neostylopyga yemenica (Bey-Bienko, 1969) comb. nov. is transferred from Duchailluia. A detailed morphological descriptions of the new species are given, N. yemenica is redescribed inclusive male genitalia. The male genitalia of N. yemenica is described for the first time. The subfamily Duchailluiinae Roth, 2003 synonymized with Blattinae Latreille, 1810.

First fossil record of Ischaliidae Blair, 1920 (Coleoptera) from Eocene Baltic amber.

Ischalia Pascoe, 1860, the monotypic genus of Ischaliidae, was placed as a subfamily among Pyrochroidae or Anthicidae by earlier authors. Nikitsky (1992) raised the status of Ischaliidae to the family rank. This position has been largely accepted and consequently followed by recent authors dealing with Ischalia, as well as in our paper. Ischaliidae is known from the Palaearctic (the Himalayas, East Asia: easternmost Russia, Japan, E China), South East Asia (southwards to Java, Borneo and the Philippines), and the Nearctic region (southern Canada, U.S.A.). No recent European representatives of Ischalia are known. At present, this genus is composed of 43 species worldwide (Gusakov & Telnov 2007; Saitô 2011; Young 2011; Young 2014; Saitô & Young 2015) placed in three subgenera (Young 2011): Ischalia s. str.-37 species, Eupleurida LeConte, 1862-5 species, Nitidischalia Young, 2011-1 species. Ischaliidae is most diverse in tropical rainforests, but many species are also known from subtropical to temperate zones. Adults and larvae of two Nearctic Ischalia feed on fungal mycelium growing on decaying wood of coniferous and broad-leaved trees (e.g. Pinus ponderosa, Lithocarpus densiflora) (Young 1985). Mimicry has been described among Bornean Ischaliidae, Tenebrionidae and Chrysomelidae (Telnov 2005) and also among Ischaliidae, Omalisidae, Lycidae and Endomychidae (Kazantsev & Young 2010).