Rare eggshell structure in Odonata: Lindenia tetraphylla (Van der Linden) (Anisoptera: Gomphidae).

Lindenia tetraphylla (Van der Linden, 1825) eggs exhibit an egg structure that is very rare in other Gomphidae species. They have a well-developed surface reticulation structure. The anterior pole of the egg has a small, rounded micropylar area consisting of seven orifices arranged radially around a central area. The posterior pole has a sessile, truncated cone that carries 55-65 coiled filaments. The filament structure found at the posterior pole of the egg has been observed in the gomphid species Lestinogomphus africanus (Fraser, 1926), Ictinogomphus australis (Selys, 1873), and I. rapax (Rambur, 1842). However, L. tetraphylla eggs differ from these species in both morphology and filament structure. This study provides a detailed analysis of the ultrastructure of L. tetraphylla eggs using scanning electron microscopy, and the functional and taxonomic significance of the eggshell are discussed. RESEARCH HIGHLIGHTS: The aim of this study is to examine the ultrastructure of the L. tetraphylla eggshell, emphasizing its function and taxonomic value. In this context, the general morphology of the egg, the reticulations on its surface, the micropylar region and micropylar structure, and the posterior filament coil were examined. In this study, the ultrastructure of L. tetraphylla egg was examined for the first time using scanning electron microscopy (SEM). As a result of the examination, it was detected for the first time that the posterior filament coil, which is rarely seen in odonate eggs, is also present in L. tetraphylla eggs. By comparing the L. tetraphylla egg with the eggs of I. ferox, I. rapax, I. australis, and L. africanus species, which are similar in terms of the posterior filament coil, the features that distinguish the L. tetraphylla egg.

Diversity of the Anisoptera & Zygoptera (Odonata: Insecta) of Swat, Pakistan / Diversidade de Anisoptera e Zygoptera (Odonata: Insecta) de Swat, Paquistão

Abstract Odonates are important biological control agents for the control of insect pests and insect disease vectors of medical and veterinary importance. The present study was conducted to evaluate the odonate fauna of Swat, Pakistan from March to October 2019. A total of 200 specimens of odonates were collected from diverse habitats. The collected specimens of the order Odonata belonged to 5 families, three families of suborder Anisoptera namely Libellulidae, Gomphidae and Aeshnidae while two families of suborder Zygoptera (Chlorocyphidae and Coenagrionidae). The specimens were categorized into 12 genera and 22 species. Libellulidae was the dominant family (n = 138) accounting for 69% of the odonate fauna. Orthetrum was the dominant genus (n = 73) of suborder Anisoptera accounting for 36.5% of the odonate fauna. The least dominant genera were Anax, Paragomphus and Rhyothemis (n = 5 each) accounting each for 2.5% of the odonate fauna. In Zygoptera, the dominant genus was Ceriagrion (12.5%) and the least dominant genus was Ischnura (6%). Pantala flavescens (Fabricius, 1798) was the most abundant odonate species in the study area recorded from all surveyed habitats. Shannon Diversity Index (H) was 2.988 and Simpson Diversity Index (D) was 0.95 for the collected odonate fauna. The highest abundance of Odonata was recorded in August, September and May while no odonate species were recorded in January, February, November and December. Lotic water bodies were the most suitable habitats with abundant odonate fauna. Anax immaculifrons (Rambur, 1842) was the largest sized odonate species having a wingspan of 53.2±1.63 mm and body length of 56.3 ± 0.4 mm. The present study shows the status of odonate fauna of Swat, Pakistan in diverse habitats and seasonsonal variation throughout the year. Further work is recommended to bridge the gaps in the existing literature.

Resumo Odonatos são importantes agentes de controle biológico para o controle de insetos-praga e vetores de doenças de insetos de importância médica e veterinária. O presente estudo foi conduzido para avaliar a fauna de odonatos de Swat, Paquistão, de março a outubro de 2019. Um total de 200 espécimes de odonatos foi coletado em diversos habitats. Os espécimes coletados da ordem Odonata pertenciam a cinco famílias, três famílias da subordem Anisoptera, a saber, Libellulidae, Gomphidae e Aeshnidae, enquanto duas famílias eram da subordem Zygoptera (Chlorocyphidae e Coenagrionidae). Os espécimes foram classificados em 12 gêneros e 22 espécies. Libellulidae foi a família dominante (n = 138), respondendo por 69% da fauna de odonatos. Orthetrum foi o gênero dominante (n = 73) da subordem Anisoptera, responsável por 36,5% da fauna de odonatos. Os gêneros menos dominantes foram Anax, Paragomphus e Rhyothemis (n = 5 cada), representando cada um 2,5% da fauna de odonatos. Em Zygoptera, o gênero dominante foi Ceriagrion (12,5%), e o gênero menos dominante foi Ischnura (6%). Pantala flavescens (Fabricius, 1798) foi a espécie de odonato mais abundante na área de estudo, registrada em todos os habitats pesquisados. O Índice de Diversidade de Shannon (H) foi de 2,988, e o Índice de Diversidade de Simpson (D) foi de 0,95 para a fauna de odonatos coletados. A maior abundância de Odonata foi registrada em agosto, setembro e maio, enquanto nenhuma espécie de Odonata foi registrada em janeiro, fevereiro, novembro e dezembro. Corpos d'água lóticos foram os habitats mais adequados, com abundante fauna de odonatos. Anax imaculifrons (Rambur, 1842) foi a espécie de odonato de maior tamanho, com envergadura de 53,2 ± 1,63 mm e comprimento do corpo de 56,3 ± 0,4 mm. O presente estudo mostrou o status da fauna de odonatos de Swat, Paquistão, em diversos habitats e variação sazonal ao longo do ano. Recomenda-se trabalho adicional para preencher as lacunas na literatura existente.

Diversity of the Anisoptera & Zygoptera (Odonata: Insecta) of Swat, Pakistan / Diversidade de Anisoptera e Zygoptera (Odonata: Insecta) de Swat, Paquistão

Odonates are important biological control agents for the control of insect pests and insect disease vectors of medical and veterinary importance. The present study was conducted to evaluate the odonate fauna of Swat, Pakistan from March to October 2019. A total of 200 specimens of odonates were collected from diverse habitats. The collected specimens of the order Odonata belonged to 5 families, three families of suborder Anisoptera namely Libellulidae, Gomphidae and Aeshnidae while two families of suborder Zygoptera (Chlorocyphidae and Coenagrionidae). The specimens were categorized into 12 genera and 22 species. Libellulidae was the dominant family (n = 138) accounting for 69% of the odonate fauna. Orthetrum was the dominant genus (n = 73) of suborder Anisoptera accounting for 36.5% of the odonate fauna. The least dominant genera were Anax, Paragomphus and Rhyothemis (n = 5 each) accounting each for 2.5% of the odonate fauna. In Zygoptera, the dominant genus was Ceriagrion (12.5%) and the least dominant genus was Ischnura (6%). Pantala flavescens (Fabricius, 1798) was the most abundant odonate species in the study area recorded from all surveyed habitats. Shannon Diversity Index (H) was 2.988 and Simpson Diversity Index (D) was 0.95 for the collected odonate fauna. The highest abundance of Odonata was recorded in August, September and May while no odonate species were recorded in January, February, November and December. Lotic water bodies were the most suitable habitats with abundant odonate fauna. Anax immaculifrons (Rambur, 1842) was the largest sized odonate species having a wingspan of 53.2±1.63 mm and body length of 56.3 ± 0.4 mm. The present study shows the status of odonate fauna of Swat, Pakistan in diverse habitats and seasonsonal variation throughout the year. Further work is recommended to bridge the gaps in the existing literature.

Odonatos são importantes agentes de controle biológico para o controle de insetos-praga e vetores de doenças de insetos de importância médica e veterinária. O presente estudo foi conduzido para avaliar a fauna de odonatos de Swat, Paquistão, de março a outubro de 2019. Um total de 200 espécimes de odonatos foi coletado em diversos habitats. Os espécimes coletados da ordem Odonata pertenciam a cinco famílias, três famílias da subordem Anisoptera, a saber, Libellulidae, Gomphidae e Aeshnidae, enquanto duas famílias eram da subordem Zygoptera (Chlorocyphidae e Coenagrionidae). Os espécimes foram classificados em 12 gêneros e 22 espécies. Libellulidae foi a família dominante (n = 138), respondendo por 69% da fauna de odonatos. Orthetrum foi o gênero dominante (n = 73) da subordem Anisoptera, responsável por 36,5% da fauna de odonatos. Os gêneros menos dominantes foram Anax, Paragomphus e Rhyothemis (n = 5 cada), representando cada um 2,5% da fauna de odonatos. Em Zygoptera, o gênero dominante foi Ceriagrion (12,5%), e o gênero menos dominante foi Ischnura (6%). Pantala flavescens (Fabricius, 1798) foi a espécie de odonato mais abundante na área de estudo, registrada em todos os habitats pesquisados. O Índice de Diversidade de Shannon (H) foi de 2,988, e o Índice de Diversidade de Simpson (D) foi de 0,95 para a fauna de odonatos coletados. A maior abundância de Odonata foi registrada em agosto, setembro e maio, enquanto nenhuma espécie de Odonata foi registrada em janeiro, fevereiro, novembro e dezembro. Corpos d’água lóticos foram os habitats mais adequados, com abundante fauna de odonatos. Anax imaculifrons (Rambur, 1842) foi a espécie de odonato de maior tamanho, com envergadura de 53,2 ± 1,63 mm e comprimento do corpo de 56,3 ± 0,4 mm. O presente estudo mostrou o status da fauna de odonatos de Swat, Paquistão, em diversos habitats e variação sazonal ao longo do ano. Recomenda-se trabalho adicional para preencher as lacunas na literatura existente.

Diversity of the Anisoptera & Zygoptera (Odonata: Insecta) of Swat, Pakistan

Abstract Odonates are important biological control agents for the control of insect pests and insect disease vectors of medical and veterinary importance. The present study was conducted to evaluate the odonate fauna of Swat, Pakistan from March to October 2019. A total of 200 specimens of odonates were collected from diverse habitats. The collected specimens of the order Odonata belonged to 5 families, three families of suborder Anisoptera namely Libellulidae, Gomphidae and Aeshnidae while two families of suborder Zygoptera (Chlorocyphidae and Coenagrionidae). The specimens were categorized into 12 genera and 22 species. Libellulidae was the dominant family (n = 138) accounting for 69% of the odonate fauna. Orthetrum was the dominant genus (n = 73) of suborder Anisoptera accounting for 36.5% of the odonate fauna. The least dominant genera were Anax, Paragomphus and Rhyothemis (n = 5 each) accounting each for 2.5% of the odonate fauna. In Zygoptera, the dominant genus was Ceriagrion (12.5%) and the least dominant genus was Ischnura (6%). Pantala flavescens (Fabricius, 1798) was the most abundant odonate species in the study area recorded from all surveyed habitats. Shannon Diversity Index (H) was 2.988 and Simpson Diversity Index (D) was 0.95 for the collected odonate fauna. The highest abundance of Odonata was recorded in August, September and May while no odonate species were recorded in January, February, November and December. Lotic water bodies were the most suitable habitats with abundant odonate fauna. Anax immaculifrons (Rambur, 1842) was the largest sized odonate species having a wingspan of 53.2±1.63 mm and body length of 56.3 ± 0.4 mm. The present study shows the status of odonate fauna of Swat, Pakistan in diverse habitats and seasonsonal variation throughout the year. Further work is recommended to bridge the gaps in the existing literature.

Resumo Odonatos são importantes agentes de controle biológico para o controle de insetos-praga e vetores de doenças de insetos de importância médica e veterinária. O presente estudo foi conduzido para avaliar a fauna de odonatos de Swat, Paquistão, de março a outubro de 2019. Um total de 200 espécimes de odonatos foi coletado em diversos habitats. Os espécimes coletados da ordem Odonata pertenciam a cinco famílias, três famílias da subordem Anisoptera, a saber, Libellulidae, Gomphidae e Aeshnidae, enquanto duas famílias eram da subordem Zygoptera (Chlorocyphidae e Coenagrionidae). Os espécimes foram classificados em 12 gêneros e 22 espécies. Libellulidae foi a família dominante (n = 138), respondendo por 69% da fauna de odonatos. Orthetrum foi o gênero dominante (n = 73) da subordem Anisoptera, responsável por 36,5% da fauna de odonatos. Os gêneros menos dominantes foram Anax, Paragomphus e Rhyothemis (n = 5 cada), representando cada um 2,5% da fauna de odonatos. Em Zygoptera, o gênero dominante foi Ceriagrion (12,5%), e o gênero menos dominante foi Ischnura (6%). Pantala flavescens (Fabricius, 1798) foi a espécie de odonato mais abundante na área de estudo, registrada em todos os habitats pesquisados. O Índice de Diversidade de Shannon (H) foi de 2,988, e o Índice de Diversidade de Simpson (D) foi de 0,95 para a fauna de odonatos coletados. A maior abundância de Odonata foi registrada em agosto, setembro e maio, enquanto nenhuma espécie de Odonata foi registrada em janeiro, fevereiro, novembro e dezembro. Corpos dágua lóticos foram os habitats mais adequados, com abundante fauna de odonatos. Anax imaculifrons (Rambur, 1842) foi a espécie de odonato de maior tamanho, com envergadura de 53,2 ± 1,63 mm e comprimento do corpo de 56,3 ± 0,4 mm. O presente estudo mostrou o status da fauna de odonatos de Swat, Paquistão, em diversos habitats e variação sazonal ao longo do ano. Recomenda-se trabalho adicional para preencher as lacunas na literatura existente.

Complete mitochondrial genome of Asiagomphus coreanus (Odonata: Gomphidae), which is endemic to South Korea.

Asiagomphus coreanus (Doi & Okumura, 1937) belongs to the family Gomphidae in the order Odonata, and has been listed as an endemic species in South Korea. Here, we assembled its complete mitochondrial genome (mitogenome) which is 15,649 base pairs (bp) in length. The A. coreanus mitogeneome consists of a typical set of genes [13 protein-coding genes (PCGs), 2 ribosomal RNA (rRNA) genes, and 22 transfer RNA (tRNA) genes] and one major non-coding A + T-rich region which is 846 bp long. The gene arrangement of the species was identical to that of commonly found in the majority of the insects. Phylogenetic analyses using the concatenated sequences of 13 PCGs and two rRNA genes of the representative odonate mitogenomes by Bayesian inference method revealed that A. coreanus belongs to the Gomphidae family with a strong nodal support (Bayesian posterior probabilities = 1). Unlike previous phylogenetic analyses (with regards to suborder relationships) the suborder Anisozygoptera-which was represented by a single species, Epiophlebia superstes-was placed as the sister to Zygoptera.

Preliminary Assessment of Chemical Elements in Sediments and Larvae of Gomphidae (Odonata) from the Blyde River of the Olifants River System, South Africa.

Benthic macroinvertebrates and sediments can act as good indicators of environmental quality. The aim of this study was to assess the accumulation of chemical elements in the Gomphidae (Odonata) collected in the Blyde River. Seven sites were sampled for river sediments assessment and five sites for larvae (naiads) of Gomphidae bioaccumulation analysis. The tissue samples were analysed using inductively coupled plasma optical emission spectrometry (ICP-OES). The results showed high levels of all of the tested elements except Cd in the sediment. The mean concentrations of As, Cu and Cr exceeded the standard guideline values, whereas Pb and Zn were below the standard guideline values. In the insect body tissue, the concentrations of most elements were higher than in the sediments. The elements with the highest concentrations were Mn, Zn, Cu, and As. The bioaccumulation factor (BF) showed a tendency for bioaccumulation for almost all of the selected elements in the insect. The BF value was high for Cu, Mn, Sb, and Zn (BF > 1). The high concentrations of elements in the insect body tissue may pose a risk to fish that consume them, and subsequently to humans when fish from the river are consumed. It is therefore important to monitor the river to reduce pollution to prevent health risks in humans, especially in communities that rely on the river for water and food.

A review of Megalogomphus sumatranus (Krüger, 1899) and its allies in Sundaland with a description of a new species from Borneo (Odonata: Anisoptera: Gomphidae).

Megalogomphus sumatranus (Krüger, 1899) and its allies in Sundaland are reviewed. The accessory genitalia of the males of this genus, hardly considered previously, are found to be taxonomically informative. The taxon from Borneo previously treated as M. sumatranus is described from both sexes as M. buddi sp. nov. (holotype ♂ Sungai Datai, Nanga Bloh, Lanjak Entimau Wildlife Sanctuary, Kapit Division, Sarawak, Malaysia, 22 viii 2013, leg. J. anak Awan M. anak Adau; deposited at the Naturalis Biodiversity Center, Leiden, the Netherlands). Megalogomphus borneensis (Laidlaw, 1914), described as a subspecies of M. icterops (Martin, 1903) and subsequently relegated to the synonymy of that species, is considered to be a distinct species. Megalogomphus icterops is however considered to be a junior synonym of M. sumatranus. A re-description of the holotype of Megalogomphus borneensis is provided as is the first description of the female. Descriptive notes with illustrations of Megalogomphus sumatranus are given.

Stylogomphus thongphaphumensis (Odonata: Anisoptera: Gomphidae), a new gomphid dragonfly and the first record of S. malayanus Sasamoto, 2001 from Thailand.

Stylogomphus thongphaphumensis sp. nov. is described from a type series of specimens reared from larvae (holotype ♂, Huai Khayeng, Thong Pha Phum district, Kanchanaburi Province; 14°36'20″N 98°34'38″E, 206 m a.s.l., larva collected on 14.XII.2014; adult emerged on 30.IV.2015). All larvae were collected from the same locality in western Thailand. Description of the larva (based on preserved exuviae) is provided as well. The adult of this species can be distinguished from other Stylogomphus Fraser, 1922, by the morphology of the male anal appendages, pterothoracic pattern, abdominal pattern, male genitalia and female valvula vulvae (detail provided in the differential diagnosis below). This is the first species of Stylogomphus to be described from Thailand. This study also reports the first record of Stylogomphus malayanus Sasamoto, 2001, from Thailand.

New synonyms and a new subspecies of Macrogomphus Selys, 1858 (Odonata: Gomphidae) from continental south-east Asia.

The taxa of the genus Macrogomphus Selys, 1858 occurring in continental south-east Asia are reconsidered. Macrogomphus rivularis Förster, 1914 (described from Vietnam), M. borikhanensis Fraser, 1933 (described from Laos), and M. guilinensis Chao, 1983 (described from China), are synonymised with M. albardae Selys, 1878. The relationship and conspecificity of the latter with M. parallelogramma Burmeister, 1839 are doubtful, perhaps they are bona species. Males of M. albardae (and seemingly of parallelogramma as well) are approximately trimorphic for the pale pattern of the abdominal S3-S6, being of either a 'dashed morph' (with small isolated anteriolateral spots and conspicuous middorsal streaks), or a 'ringed morph (with broad anterior rings and less conspicuous middorsal streaks), or an 'intermediate morph'. Validity of the species M. matsukii Asahina, 1986 is doubted; its holotype could be an aberrant male of M. albardae. The main diagnostic character of M. albardae, M. phalantus Lieftinck, 1935 and probably M. parallelogramma, is clarified to be the structure of the cercus inner branch. M. phalantus jayavarman subsp. nov. is described from temporarily inundated forest at the northern bank of the great Lake Tonlé Sap of Cambodia (Siem Reap Province, 1.5 km SSW of Kampong Pluk village, 13.1956° N, 103.9725° E, 3 m a.s.l.), which is ca 1,300 km north and overseas from the presumed range of M. phalantus phalantus.

A new species of Cyclogomphus Selys, 1854 (Insecta: Odonata: Gomphidae) from the Western Ghats, India with comments on the status of Cyclogomphus vesiculosus Selys, 1873.

A new species of Cyclogomphus Selys, 1854 is described from the Western Ghats of India based on male and female specimens collected from Goa and Kerala states. The new species differs from all the known species of Cyclogomphus in the shape of the anal appendages as well as the thoracic and abdominal markings. We also provide here an updated key to the Cyclogomphus species and comment on the status of Cyclogomphus vesiculosus Selys, 1873.

A new species of Gomphidia Selys, 1854 (Insecta: Odonata: Anisoptera: Gomphidae) from the Western Ghats of India.

A new species of Gomphidia Selys, 1854 is described from southern Western Ghats, India based on a male specimen. The new species is very distinct from all the known species of Gomphidia from Indian subcontinent. Revised keys to the males and known females of Gomphidia of India and Sri Lanka are provided.

A new species of Heliogomphus from Vietnam (Odonata: Gomphidae).

Heliogomphus bidentatus sp. nov. (holotype male: Tam Dao National Park, Vinh Phuc Prov., northern Vietnam) is described from north and central Vietnam. This new species is similar to H. scorpio (Ris, 1912), but can be separated by the coloration of the thorax and details of the male caudal appendages. Information on its biology and ecology is provided.

Description of a female and variation of Microgomphus alani Kosterin, 2016 (Odonata: Gomphidae) in Cambodia, with a note on sexual dimorphism in Microgomphus spp.

Microgomphus alani Kosterin, 2016 was described from two teneral males from the Cardamom Mts. in SW Cambodia; it turned out that the description was based on the holotype with the anal appendages not fully expanded. Further specimens of M. alani (all mature) are reported: a male and two females from the Phnom Kulen Mts, NW Cambodia and five males and one female from Mondulkiri Province, E Cambodia. The appendage shape in mature males is illustrated and discussed and the female is described for the first time. The specimens from E Cambodia are ca 15-20% larger than those from SW and NW Cambodia and exhibit some differences in the shapes of the male epiproct and anterior hamulus and face maculation, however evaluated as insufficient to erect a new taxon. A hitherto neglected manifestation of sexual dimorphism in probably most Microgomphus spp. and at least some Heliogomphus spp. is pointed out. It concerns the mid- and metafemur, in males armed with numerous, dense and small spinulets but with two rows of long, sparse spines in females. An earlier report of Microgomphus jurzitzai Karube, 2000 for Cambodia was based on a misidentified M. alani specimen and is cancelled. A female specimen earlier identified and described from Thailand as Heliogomphus selysi Fraser, 1925 most probably was M. alani as well.

Description of Microgomphus farrelli sp. nov. (Odonata: Anisoptera: Gomphidae) based on adults of both sexes and larvae from Northern Thailand.

The new gomphid species, Microgomphus farrelli sp. nov., is described and illustrated on the basis of male and female adult specimens and larvae collected from Chiang Mai and Mae Hong Son province, Northern Thailand. It is compared with other species of the genus. Based on the larvae this species is most closely related to Microgomphus svihleri (Asahina, 1970), comb. nov., which is the senior and valid synonym of Microgomphus thailandicus Asahina, 1981, syn. nov.

The genus Asiagomphus in Vietnam, with descriptions of three new species, and first descriptions of the male of Asiagomphus auricolor (Fraser, 1926) and of the female of Asiagomphus reinhardti Kosterin Yokoi, 2016 (Odonata: Gomphidae).

The occurrence in Vietnam of eight species of Asiagomphus Asahina, 1985 is discussed. Three new species are described (Asiagomphus kosterini [holotype: Da Lat, Lam Dong Prov., (appr. 11.880 N, 108.442 E), 17-IV-2016], Asiagomphus monticola [holotype: Xuan Son NP, Phu Tho Prov., (appr. 21.163 N, 104.896 E), 31-V-2014], and Asiagomphus superciliaris [holotype: Huu Lien NR, Lang Son Prov., (appr. 21.686 N, 106.376 E), 01-V-2014], and the male of Asiagomphus auricolor (Fraser, 1926) and the female of Asiagomphus reinhardti Kosterin Yokoi, 2016 are described for the first time. The female of Asiagomphus auricolor is redescribed. Distribution and ecology of all species and a key to the males occurring in Vietnam are provided.

Anormogomphus kiritshenkoi Bartenev, 1913 (Odonata: Gomphidae): a literature review of the variable spelling of the species epithet, choice of the correct spelling and notes on the type locality of the species.

The spelling of the specific name of an Anormogomphus species in its original description by Bartenev (1913) was variable, kiritshenkoi (5 times) vs kiritschenkoi (1 time). Bartenev himself did not mention this species in his further publications. Later authors proceeded to use different spelling versions of this name, including those not in the original paper. According to the International Code of Zoological Nomenclature (ICZN), the valid spelling should be chosen by the First Reviser. Acting as such, we choose as valid the name spelling Anormogomphus kiritshenkoi Bartenev, 1913, since it predominates in the original description, appears in the species subtitle, and is a proper transliteration from Cyrillic to Latin. It also corresponds to the spelling of his name used by A.N. Kiritshenko himself. Information of the type locality of the species is provided.

Revision of the Genus Leptogomphus Selys in Borneo, including gene trees and a two marker molecular phylogeny (Odonata: Anisoptera: Gomphidae).

The Bornean members of the genus Leptogomphus Selys are revised. Two new species are described: Leptogomphus schieli sp. nov. (holotype ♂, Gunung Penrissen, Kuching Division, Sarawak, Malaysia, to be deposited in BMNH) and Leptogomphus sii sp. nov. (holotype ♂, Sungai Sii, upper Baram, Miri Division, Sarawak, Malaysia, in RMNH). Leptogomphus mariae Lieftinck, 1948 is considered to be a junior synonym of L. coomansi Laidlaw, 1936. The true male of L. pasia van Tol, 1990 is described for the first time; male specimens previously treated as L. pasia or L. cf pasia actually belong to a taxon closely allied to, and possibly merely a form of, L. coomansi. A description is given of the female of another new species, but the species is not named in the absence of the male. Female specimens from south-western Sarawak, similar to L. williamsoni Laidlaw, 1912, are considered likely to also represent a distinct species. The female of L. pendleburyi Laidlaw, 1934 is described for the first time and fresh descriptions of the males of L. coomansi, L. pendleburyi and L. williamsoni, and the female of L. coomansi are given. Keys to both sexes, and distribution maps are given. A molecular analysis of the Bornean species (except L. schieli) using the COI and ITS markers is presented.

A new species of the genus Burmagomphus Williamson (Odonata: Gomphidae) from Northern Thailand.

Burmagomphus chiangmaiensis sp. nov. (holotype: Ban Luang, Chom Thong, Chiang Mai province, Thailand, 890-900 m, 14 v 2012) is described and illustrated. It can be differentiated from its most similar congener, B. apricus from China, by shape of posterior hamulus, yellow trapezoid band on occiput, and larger size.

Two new dragonflies (Odonata: Anisoptera) from the Miocene of Carinthia (Austria).

Two new species of fossil dragonflies from the Middle Miocene fossil site Schaßbach (Carinthia, Austria) are described. The presence of Gomphaeschna carinthiae sp. nov. and Ictinogomphus hassleri sp. nov. in the fossil record of Central Europe confirms the scenario of a more widespread distribution of the represented genera in the Miocene in contrast to their Recent distribution.

The larvae of Epigomphus jannyae Belle, 1993 and E. tumefactus Calvert, 1903 (Insecta: Odonata: Gomphidae).

The taxonomic knowledge about immature stages of the insect order Odonata (dragonflies and damselflies) is rather limited in tropical America. Here, the larvae of Epigomphus jannyae Belle, 1993 and E. tumefactus Calvert, 1903 are described, figured, and compared with other described congeners. E. jannyae larva is characterized by 3rd antennomere 1.6 times longer than its widest part; ligula very poorly developed, with ten short, truncate teeth on middle; apical lobe of labial palp rounded and smooth. Lateral margins on abdominal segments (S5-9) serrated, lateral spines on S6-9 small and divergent; male epiproct with a pair of dorsal tubercles at basal 0.66; tips of cerci and paraprocts strongly divergent. The larva of E. tumefactus is characterized by 3rd antennomere 2.3 times longer than its widest part, ligula with 6-7 truncate teeth, apical lobe of labial palp acute and finely serrate. Lateral margins of S6-9 serrate, lateral spines on S7-9; male epiproct with a pair of dorsal tubercles at basal 0.50. Differences with other species were found in 3rd antennomere, lateral spines of S7-9, and the caudal appendages. Epigomphus larvae inhabit small, shallow creeks (1st order streams) where they live in fine benthic sediments. When mature, the larva leaves the water in shady places, climbing small rocks at the water's edge and metamorphosing horizontally on flat rocks. These new descriptions bring the total number of Epigomphus species with known larval stages to eight; only 28% of the species in this genus are known as larva.