The earliest Ethiopian wolf: implications for the species evolution and its future survival.

In 2017, a hemimandible (MW5-B208), corresponding to the Ethiopian wolf (Canis simensis), was found in a stratigraphically-controlled and radio-isotopically-dated sequence of the Melka Wakena paleoanthropological site-complex, on the Southeastern Ethiopian Highlands, ~ 2300 m above sea level. The specimen is the first and unique Pleistocene fossil of this species. Our data provide an unambiguous minimum age of 1.6-1.4 Ma for the species' presence in Africa and constitutes the first empirical evidence that supports molecular interpretations. Currently, C. simensis is one of the most endangered carnivore species of Africa. Bioclimate niche modeling applied to the time frame indicated by the fossil suggests that the lineage of the Ethiopian wolf faced severe survival challenges in the past, with consecutive drastic geographic range contractions during warmer periods. These models help to describe future scenarios for the survival of the species. Projections ranging from most pessimistic to most optimistic future climatic scenarios indicate significant reduction of the already-deteriorating territories suitable for the Ethiopian Wolf, increasing the threat to the specie's future survival. Additionally, the recovery of the Melka Wakena fossil underscores the importance of work outside the East African Rift System in research of early human origins and associated biodiversity on the African continent.

Artifact3-D: New software for accurate, objective and efficient 3D analysis and documentation of archaeological artifacts.

The study of artifacts is fundamental to archaeological research. The features of individual artifacts are recorded, analyzed, and compared within and between contextual assemblages. Here we present and make available for academic-use Artifact3-D, a new software package comprised of a suite of analysis and documentation procedures for archaeological artifacts. We introduce it here, alongside real archaeological case studies to demonstrate its utility. Artifact3-D equips its users with a range of computational functions for accurate measurements, including orthogonal distances, surface area, volume, CoM, edge angles, asymmetry, and scar attributes. Metrics and figures for each of these measurements are easily exported for the purposes of further analysis and illustration. We test these functions on a range of real archaeological case studies pertaining to tool functionality, technological organization, manufacturing traditions, knapping techniques, and knapper skill. Here we focus on lithic artifacts, but the Artifact3-D software can be used on any artifact type to address the needs of modern archaeology. Computational methods are increasingly becoming entwined in the excavation, documentation, analysis, database creation, and publication of archaeological research. Artifact3-D offers functions to address every stage of this workflow. It equips the user with the requisite toolkit for archaeological research that is accurate, objective, repeatable and efficient. This program will help archaeological research deal with the abundant material found during excavations and will open new horizons in research trajectories.

Kaizer Hill (Modi'in), a pre-pottery neolithic a quarry site - the terraced slopes.

The research of the Kaizer Hill site (the Hilltop and its Terraces), recognized as a Pre Pottery Neolithic A (PPNA) quarry site, involved studies of the rock damage associated with the quarrying activities as well as of the recovered material remains, mostly chipped stone artifacts. We present here the results of our on-site explorations (excavations, surveys and surface-collections), focusing on the findings deriving from the Terraces. Diverse rock damage patterns were identified and described, portraying systematic rock mass-exploitation through quarrying fronts, natural rock joints and fissures enlargement, drilling and chiseling. There are multiple indications that the local bedrock (Bi'na Formation, Turonian) comprising flint and limestone was quarried under a systematic quality evaluation, leaving residual flint unsuitable for exploitation. Of interest to note that nearly all of the flint artifacts excavated and collected on the Terraces were made on raw material transported from the Hilltop (Mishash Formation, Campanian), knapped in-situ, on the quarried rock surfaces of the slopes. The flint tools bear witness to intensive use involving mainly boring and drilling. The dominant tool type is the flint axe for which a variety of waste products related to its production were found in-situ, enabling the reconstruction of axe reduction sequence. Similar axes and waste products were found in many PPN sites indicating that there was a common, widely-used scheme of making flint axes during the PPN. Interestingly, besides the flint waste, there were also limestone waste products typical of the last shaping and thinning stages of axe production, indicating that limestone axes were shaped technologically similar to the flint ones, contrary to what has been assumed before. Rare findings, such as obsidian pieces, originating from much further a-field indicate ties with other PPN communities, near and/or far. Overall, this study provides unique and novel insights on Levantine PPN lifeways.

Do a few tools necessarily mean a few people? A techno-morphological approach to the question of group size at Gesher Benot Ya'aqov, Israel.

The question of Paleolithic group size has been addressed by scholars in many disciplines applying different methods. In our study we apply a novel analytical approach in an attempt to assess the group size of hominins that occupied the Acheulian site of Gesher Benot Ya'aqov, Israel (GBY). Within this framework, we subjected the handaxe assemblages from several archaeological horizons at the site to a morpho-technological analysis. The analysis combined high-resolution three-dimensional geometric morphometric analysis with typo-technological attribute analysis to assess the inter- and intra-assemblage morpho-technological variability. The analysis was also applied to an experimental handaxe assemblage produced by an expert knapper. The results of the analysis show high morphological homogeneity coupled with high technological variability in each of the archaeological assemblages. This pattern is highly indicative of the work of expert knappers, as is also suggested by the comparison between the archaeological and experimental assemblages. The high density of archaeological remains in some of the GBY occupations and their pristine taphonomic condition provide additional support for the involvement of large groups of hominins, although some horizons are far poorer in archaeological remains and hence do not allow such an interpretation. Nevertheless, the fact that in all assemblages the handaxes show the same techno-morphological pattern indicates that they were all produced by expert knappers. As shown by numerous models and ethnographic data, the presence of experts can be viewed as an indication of large and socially complex societies. Thus, although some of the GBY occupations were not formed by large groups, the smaller groups whose activities are recorded were very likely to be part of larger, socially complex cultural groups. This variability in occupational intensity is interpreted as representing an aggregation-dispersal mechanism, similar to those documented in many hunter-gatherer societies.

AGMT3-D: A software for 3-D landmarks-based geometric morphometric shape analysis of archaeological artifacts.

We present here a newly developed software package named Artifact GeoMorph Toolbox 3-D (AGMT3-D). It is intended to provide archaeologists with a simple and easy-to-use tool for performing 3-D landmarks-based geometric morphometric shape analysis on 3-D digital models of archaeological artifacts. It requires no prior knowledge of programming or proficiency in statistics. AGMT3-D consists of a data-acquisition procedure for automatically positioning 3-D models in space and fitting them with grids of 3-D semi-landmarks. It also provides a number of analytical tools and procedures that allow the processing and statistical analysis of the data, including generalized Procrustes analysis, principal component analysis, a warp tool, automatic calculation of shape variabilities and statistical tests. It provides an output of quantitative, objective and reproducible results in numerical, textual and graphic formats. These can be used to answer archaeologically significant questions relating to morphologies and morphological variabilities in artifact assemblages. Following the presentation of the software and its functions, we apply it to a case study addressing the effects of different types of raw material on the morphologies and morphological variabilities present in an experimentally produced Acheulian handaxe assemblage. The results show that there are statistically significant differences between the mean shapes and shape variabilities of handaxes produced on flint and those produced on basalt. With AGMT3-D, users can analyze artifact assemblages and address questions that are deducible from the morphologies and morphological variabilities of material culture assemblages. These questions can relate to issues of, among others, relative chronology, cultural affinities, tool function and production technology. AGMT3-D is aimed at making 3-D landmarks-based geometric morphometric shape analysis more accessible to archaeologists, in the hope that this method will become a tool commonly used by archaeologists.

Expert cognition in the production sequence of Acheulian cleavers at Gesher Benot Ya'aqov, Israel: A lithic and cognitive analysis.

Stone cleavers are one of the most distinctive components of the Acheulian toolkit. These tools were produced as part of a long and complex reduction sequence and they provide indications for planning and remarkable knapping skill. These aspects hold implications regarding the cognitive complexity and abilities of their makers and users. In this study we have analyzed a cleaver assemblage originating from the Acheulian site of Gesher Benot Ya'aqov, Israel, to provide a reconstruction of the chaîne opératoire which structured their production. This reduction sequence was taken as the basis for a cognitive analysis which allowed us to draw conclusion regarding numerous behavioral and cognitive aspects of the GBY hominins. The results indicate that the cleavers production incorporated a highly specific sequence of decisions and actions which resulted in three distinct modes of cleavers modification. Furthermore, the decision to produce a cleaver must have been taken very early in the sequence, thus differentiating its production from that of handaxes. The substantial predetermination and the specific reduction sequence provide evidence that the Gesher Benot Ya'aqov hominins had a number of cognitive categories such as a general 'tool concept' and a more specific 'cleaver concept', setting them apart from earlier tool-producing hominins and extant tool-using non-human primates. Furthermore, it appears that the Gesher Benot Ya'aqov lithic technology was governed by expert cognition, which is the kind of thinking typical of modern human experts in their various domains. Thus, the results provide direct indications that important components of modern cognition have been well established in the minds of the Gesher Benot Ya'aqov hominins.

A new type of anvil in the Acheulian of Gesher Benot Ya'aqov, Israel.

We report here on the identification and characterization of thin basalt anvils, a newly discovered component of the Acheulian lithic inventory of Gesher Benot Ya'aqov (GBY). These tools are an addition to the array of percussive tools (percussors, pitted stones and anvils) made of basalt, flint and limestone. The thin anvils were selected from particularly compact, horizontally fissured zones of basalt flows. This type of fissuring produces a natural geometry of thick and thin slabs. Hominins at GBY had multiple acquisition strategies, including the selection of thick slabs for the production of giant cores and cobbles for percussors. The selection of thin slabs was carried out according to yet another independent and targeted plan. The thinness of the anvils dictated a particular range of functions. The use of the anvils is well documented on their surfaces and edges. Two main types of damage are identified: those resulting from activities carried out on the surfaces of the anvils and those resulting from unintentional forceful blows (accidents de travaille). Percussive activities that may have been associated with the thin anvils include nut cracking and the processing of meat and bones, as well as plants.