A Rare Cleptoparasitic Bee Tetralonioidella himalayana (Bingham, 1897) (Hymenoptera: Apidae) from India: Review and New Data.

Tetralonioidella Strand 1914 is a very rare apid genus globally, of which only two species, Tetralonioidella himalayana (Bingham, 1897) and Tetralonioidella tricolor (Lieftinck, 1972), are known from India. They are known to be cleptoparasitic on species of Habropoda Smith, 1854 and Elaphropoda Lieftinck, 1966. During hymenopteran survey in Arunachal Pradesh, we encountered Tetralonioidella himalayana (Bingham, 1897) and observed their behavioral regime, which is very poorly known. The identifying characteristics of both male and female, their foraging behavior, floral preference, distribution pattern, and possible host association have been studied. Exclusive floral association and host specialization are potential contributing factors to the rarity of T. himalayana. Such factors may limit the distribution range of the species. A priori sampling resolution with genetic and demographic exploration is required to evaluate the present status of such bee species.

Two new species and two new records of the genus Tenuibaetis Kang & Yang, 1994 (Ephemeroptera: Baetidae) from Indian Himalaya.

As part of ongoing exploration of the mayflies of streams and rivers of Indian Himalayan regions, we describe two new species of the genus Tenuibaetis based on recently collected materials from Western Himalaya viz., Tenuibaetis himani Kubendran, Vasanth & Subramanian sp. nov. and Tenuibaetis kangi Kubendran, Vasanth & Subramanian sp. nov. Additionally, two new records from Eastern Himalaya viz., T. arduus (Kang & Yang, 1994) and T. inornatus (Kang & Yang, 1994) are presented. The biological notes of the stream/rivers are appended. A key to the known Indian species of the genus Tenuibaetis and a distribution maps are provided.

A review of the Drepanosticta carmichaeli-group, with the description of D. wildermuthi sp. nov. from the Central Highlands of Vietnam (Odonata: Zygoptera: Platystictidae).

A modified and expanded definition of the Drepanosticta carmichaeli-group is given. This includes the species: D. annandalei Fraser, 1924, D. brownelli Tinkham, 1938, D. carmichaeli (Laidlaw, 1915), D. emtrai Dow, Kompier Phan, 2018, D. hongkongensis Wilson, 1997, D. jurzitzai Hmlinen, 1999, D. sumatrana Sasamoto Karube, 2007, D. tenella Lieftinck, 1935, D. vietnamica Asahina, 1997 and D. wildermuthi sp. nov.). The species of the group are reviewed and in most cases illustrated, diagnostic notes are given wherever possible. Drepanosticta polychromatica Fraser, 1931 is considered to be a junior synonym of D. carmichaeli and variation in D. carmichaeli is discussed. Drepanosticta wildermuthi sp. nov. from the Central Highlands of Vietnam (holotype male from Bao Loc district, Lam Dong Province) is described. The female of D. jurzitzai Hmlinen, 1999 is described for the first time. A key to the males of the Drepanosticta carmichaeli-group is provided.

A new species and new record of the genus Platybaetis Mller-Liebenau, 1980 (Ephemeroptera: Baetidae) from Eastern Himalaya, India.

A new species of Platybaetis viz., P. selvai sp. nov. is described herein based on larval collections from Tangon stream in the state of Arunachal Pradesh in Eastern region of Indian Himalaya. It can be differentiated by the following combination of characters: (i) posterior margin of abdominal segments IX with rounded 'U shaped spines; (ii) anterolateral margin of gills IVII with minute setae; (iii) claw with 78 denticles; (iv) paracercus composed of 1011 segments; (v) hindwing pads reduced, small. Brief ecological notes are appended.

Overview of Indian and Nepali representatives of the Cincticostella nigra (Uéno, 1928) complex (Ephemeroptera, Ephemerellidae), with discussion about Cincticostella Allen, 1971 species complexes.

The concept of the Cincticostella nigra (Uéno, 1928) (Ephemeroptera: Ephemerellidae) complex is clarified and Indian and Nepali representatives of the complex are reviewed. Four new species are described viz. Cincticostella changfai Martynov & Palatov, sp. nov., Cincticostella funki Martynov, Selvakumar, Palatov & Vasanth, sp. nov., Cincticostella shinichii Martynov & Palatov, sp. nov. and Cincticostella wangi Selvakumar, Martynov & Subramanian, sp. nov. The larva of C. corpulenta (Braasch, 1981) is re-described, based on the holotype and paratypes. Cincticostella gosei (Allen, 1975) is recorded from India for the first time. Morphological differences of the species complexes of Cincticostella Allen, 1971 are analysed. A new species complex, Cincticostella gosei complex, is proposed. Summaries of distribution data and habitat preferences, as well as new larval diagnoses, are presented for all species of the Cincticostella nigra complex.

Contribution to the study of Epeorus Eaton, 1881 (Ephemeroptera: Heptageniidae) from India.

Seven species of the genus Epeorus Eaton, 1881 are described so far from India of which redescription is attempted herein for 3 species viz. E. (C.) psi Eaton 1883, E. (E.) gilliesi Braasch 1981, E. (E.) petersi Sivaruban, Venkataraman Sivaramakrishnan 2013 based on the fresh collections. Four species viz., E. (E.) aculeatus Braasch 1990, E. (E.) bifurcatus Braasch Soldán 1979, E. (E.) unicornutus Braasch 2006 and E. (E.) unispinosus Braasch 1980 are new records to this country. Totally, 11 species of the genus Epeorus from India are dealt with herein of which six belongs to the subgenus Epeorus viz. E. (E.) aculeatus Braasch 1990, E. (E.) bifurcatus Braasch Soldán 1979, E. (E.) gilliesi Braasch, 1981 and E. (E.) petersi Sivaruban, Venkataraman Sivaramakrishnan 2013, E. (E.) unispinosus Braasch 1980, E. (E.) unicornutus Braasch 2006 and four to the subgenus Caucasiron viz., E. (C.) kapurkripalanorum Braasch 1983, E. (C.) papillatus Braasch, 2006a, E. (C.) psi Eaton 1883 which larva is described in details for the first time and E. (C.) suspicatus Braasch, 2006b . The subgeneric status of E. lahaulensis Kapur Kripalani 1963 cannot be fixed due to non-availability of larval description. A dichotomous key to the larvae of available species in India is presented. Distribution of subgenera and species within India is discussed.

Overview of Indian Hyrtanellini (Ephemeroptera: Ephemerellidae), with new species and records from related regions.

The Hyrtanellini (Ephemeroptera: Ephemerellidae: Ephemerellinae) from India and related regions are reviewed, including three new species viz., Serratella palatovi Martynov, Selvakumar Jacobus, sp. nov., Torleya simbalbarensis Selvakumar, Subramanian, Martynov Jacobus, sp. nov., and Torleya dibruensis Selvakumar, Martynov Jacobus, sp. nov.; and first country records of Teloganopsis jinghongensis (Xu, You Hsu, 1984) and Hyrtanella grandipennis (Zhou, Su Gui, 2000). Description of Serratella palatovi sp. nov. is based on material from Thailand, India and Nepal; Torleya simbalbarensis sp. nov. and Torleya dibruensis sp. nov. are described based on material from India. For now, the tribe Hyrtanellini in India comprises eight species from the genera Serratella Edmunds, 1959, Torleya Lestage, 1917, Teloganopsis Ulmer, 1939 and Hyrtanella Allen Edmunds, 1976. Additional records of Teloganopsis jinghongensis from Thailand, Torleya nepalica (Allen Edmunds, 1963) and Torleya coheri (Allen Edmunds, 1963) from Nepal also are provided here. All Indomalayan Hyrtanellini species are discussed to some extent.

New record of Megalestes gyalsey Gyeltshen, Kalkman amp; Orr, 2017 (Zygoptera: Synlestidae) from India, with first description of female and larva.

Megalestes gyalsey Gyeltshen, Kalkman Orr, 2017 is recorded for the first time from India, extending the known geographic range of the species. This report is based on the collection of 5 individuals (4 males, 1 female) from Jang waterfall, Tawang, Arunachal Pradesh and 2 males from Neora Valley National Park, Kalimpong district, West Bengal. The female of M. gyalsey is described for the first time with notes on the variation in the male. A probable larva of the species is also described and illustrated.

Three new species of Atalophlebiinae (Ephemeroptera: Leptophlebiidae) of India, with a new record of the genus Megaglena Peters and Edmunds, 1970.

Three new species of Leptophlebiidae viz., Choroterpes (Choroterpes) andamanensis Vasanth, Subramanian Selvakumar n. sp., Edmundsula meghamalaiensis Vasanth, Subramanian Selvakumar n. sp. and Megaglena agasthiya Vasanth, Subramanian Selvakumar n. sp. are described based on our recent exploration of mayflies from the streams of Andaman Island, Tamil Nadu and Kerala part of the southern Western Ghats. The genus Megaglena Peters Edmunds, 1970 has extended the distribution in southern peninsular India as represented by the occurrence of M. agasthiya n. sp. Larval key is provided to the known and new species of subgenus Choroterpes of India along with the distribution map of new species.

A new species of Notacanthurus Tshernova, 1974 (Ephemeroptera: Heptageniidae) from India.

A new species of Notacanthurus Tshernova, 1974 is described based on larvae from India. Notacanthurus pange Vasanth, Selvakumar Subramanian sp. nov. can be distinguished from all known species of Notacanthurus by the following combination of characters: (i) gill lamellae 5-6 with accessory lobe; (ii) dorsal surface of femora with scattered spatulate setae (iii) male genitalia having penis lobes fused, 'V' shaped apically blunt with me-dian pair of spine-like titillators subapically and mesally; (iv) labrum with anterior portion straight and laterally curved; and (v) lingua of hypopharynx apically broad and convex. The distribution map of the new species is also provided.

Orthetrum erythronigrum sp. nov. (Odonata: Libellulidae) from the Great Nicobar Island, India.

The male and female of Orthetrum erythronigrum sp. nov. are described from Great Nicobar Island and are believed to be endemic to the Nicobar Islands archipelago (holotype ♂, India, Andaman and Nicobar Islands, Great Nicobar Island, Great Nicobar Biosphere Reserve, N 6.99067, E 93.871363; 01-xii-2018; deposited in ZSI, SRC, Chennai, India). The male of this new species is easily distinguished from other known Orthetrum species by its black abdomen with contrasting crimson belly. Female is distinguished by its large size, the black non-metallic head, black thorax and the red abdomen with a distinct black pattern. Based on field observations and photographs, notes on the life colouration of the mature male and information on the distribution and habitat of this Great Nicobar endemic are provided.

A new species of Orthetrum Newman, 1833 (Odonata: Libellulidae) from the Andaman Islands, India.

Orthetrum andamanicum sp. nov. (holotype ♂: India, Andaman and Nicobar Islands, South Andaman island, Chidiyatapu, N 11.51, E 92.71; 08-xii-1998; deposited in RMNH, Leiden; RMNH.INS.1152911), is described as new to science. Based on additional photographic records, notes on mature males' life colouration, distribution and habitat of this Andaman endemic are provided.

Three new species of Protosticta Selys, 1885 (Odonata: Zygoptera: Platystictidae) from the Western Ghats, India, with taxonomic notes on P. mortoni Fraser, 1922 and rediscovery of P. rufostigma Kimmins, 1958.

Three new species of Protosticta Selys, 1885 (Odonata: Zygoptera: Platystictidae) from the Western Ghats biodiversity hotspot in India: P. cyanofemora sp. nov. (wet evergreen forests in Shendurney Wildlife Sanctuary, Kollam, Kerala and Kalakkad Mundanthurai Tiger Reserve, Tirunelveli, Tamil Nadu), P. myristicaensis sp. nov. (Myristica swamp at Kathalekan, Shivamogga, Karnataka) and P. sholai sp. nov. (montane sholas of Upper Manalar, Meghamalai Wildlife Sanctuary, Theni, Tamil Nadu) are described and illustrated. We compare these three new species with other Protosticta spp. from the Western Ghats based on new material and provide comprehensive differential diagnoses with determination key for males of all species occurring in the Western Ghats. The taxonomic validity of P. mortoni Fraser, 1922 is established with fresh specimens from Hassan, Karnataka, and rediscovery of P. rufostigma Kimmins, 1958 is reported from Kanyakumari Wildlife Sanctuary, Tamil Nadu.

Checklist of the dragonflies and damselflies (Insecta: Odonata) of Bangladesh, Bhutan, India, Nepal, Pakistan and Sri Lanka.

A checklist of the dragonflies and damselflies occurring in Bangladesh, Bhutan, India (including Andaman and Nicobar Islands), Nepal, Pakistan and Sri Lanka is presented. In total 588 (including 559 full species) taxa are known to occur in the region of which 251 taxa (species subspecies) are single country endemics. Recent taxonomic changes relevant to the area are summarized. Sixteen taxa are synonymized and a checklist of all synonyms established since 1950 is provided. Information is given on available larval descriptions including a list of genera present in the region for which no larvae have yet been described. Numerous species occurring in the area are still poorly known and a list of genera for which a revision is urgently needed is provided.                The following new synonyms are established: Calicnemia sudhaae Mitra, 1994 = Calicnemia imitans Lieftinck, 1948 syn. nov.; Ceriagrion fallax cerinomelas Lieftinck, 1927 = Ceriagrion fallax Ris, 1914 syn. nov.; Ceriagrion fallax pendleburyi Laidlaw, 1931 = Ceriagrion fallax Ris, 1914 syn. nov.; Coenagrion kashmirus Chowdhary Das, 1975 = Ischnura forcipata Morton, 1907 syn. nov.; Enallagma insula Fraser, 1920 = Aciagrion occidentale Laidlaw, 1919 syn. nov.; Himalagrion pithoragarhicum Sahni, 1964 = Ceriagrion fallax Ris, 1914 syn. nov.; Ischnura bhimtalensis Sahni, 1965 = Ischnura rubilio Selys, 1876 syn. nov.; Onychargia indica Sahni, 1964 = Paracercion calamorum (Ris, 1916) syn. nov.; Anaciaeschna kashmirense Singh Baijal, 1954 = Anaciaeschna martini (Selys, 1897) syn. nov.; Cyclogomphus vesiculosus Selys, 1854 = Cyclogomphus ypsilon Selys, 1954 syn. nov.; Chlorogomphus brittoi Navás, 1934 = Chlorogomphus xanthoptera (Fraser, 1919) syn. nov.; Hylaeothemis indica Fraser, 1946 = Hylaeothemis apicalis Fraser, 1924 syn. nov.; Sympetrum durum Bartenef, 1916 = Sympetrum striolatum commixtum Selys, 1884 syn. nov.; Sympetrum himalayanum Navás, 1934 = Sympetrum hypomelas (Selys, 1884) syn. nov.; Sympetrum haematoneura Fraser, 1924 = Sympetrum speciosum Oguma, 1915 syn. nov.; Sympetrum speciosum taiwanum Asahina, 1951 = Sympetrum speciosum Oguma, 1915 syn. nov.                In addition, Periaeschna lebasi Navás, 1930 is regarded a nomen nudum. The following new combinations are proposed: Onychogomphus acinaces Laidlaw, 1922 is transferred to the genus Melligomphus Chao, 1990 resulting in Melligomphus acinaces (Laidlaw, 1922) comb. nov. Onychogomphus saundersii Selys, 1854 is transferred to the genus Nychogomphus Carle, 1986 resulting in Nychogomphus saundersii (Selys, 1854) comb. nov..

New record of the genus Baetiella Uéno, 1931 (Ephemeroptera: Baetidae) from India with description of a new species and new records for five species.

A new species of the genus Baetiella is described based on larvae collected from Arunachal Pradesh, India. Baetiella subansiri Vasanth, Selvakumar Subramanian n. sp. can be distinguished by the following combination of characters: (i) gills (or gill sockets) on segments I-VI; gills I-V elongate, without tracheation, with numerous scattered pores on the surface and smooth fine simple setae along margin; gill VI reduced; (ii) posterior margin of metanotum and abdominal terga I-II each with a single posteromedian protuberance, terga III-IX each with a pair of much longer protuberances; and (iii) claw with two rows of denticles, outer row with 6-7 acute denticles increasing in length apically, inner row with 6-7 short and blunt denticles medially, subequal in length. New records of Baetiella armata Braasch, 1983, B. ausobskyi Braasch, 1983, B. imanishii Braasch, 1983, B. marginata Braasch, 1983 and B. spathae Shi Tong, 2015 from Arunachal Pradesh, India are presented based on the collection of larvae. The four species of Baetiella described by Braasch (1983) are redescribed based on freshly collected material. The Known distribution of genus Baetiella from Arunachal Pradesh, India is also provided.

A new record of the family Isonychiidae (Insecta: Ephemeroptera) from the Western Ghats, India with a description of new species.

A new species belonging to the subgenus Isonychia, of the genus Isonychia Eaton, 1871, is described based on larvae and imagoes collected from Moyar River, Nilgiri District, Tamil Nadu, India. The imagoes of I. moyarensis n. sp. can be distinguished from other described Oriental species of Isonychia (Isonychia) by the combination of characters: (i) forewing with rusty brown maculae in the costal, subcostal, and median areas; (ii) femur and tibia brown, fore leg pale, apices of tarsal segments brownish; (iii) males with distal angles of penes rounded, without serrations; (iv) second segment of gonostylus uniformly convex; and (v) sterna of tenth abdominal segment in female deeply cleft. Isonychia (Isonychia) moyarensis n. sp. can be distinguished in the larval stage from other known Oriental species by the following combination of characters: (i) abdominal terga II-IX with median dark brown maculae progressively larger with dark brown slanting streaks in lateral margins; (ii) trachea of abdominal gills I-VII unbranched; (iii) posterolateral projections on abdominal segments I-VII blunt and progressively longer than those of segments VIII-IX, sharp and distinct; and (iv) abdominal terga X pale yellow in anterior ⅓, and dark brown in the posterior ⅔. A key to the known larvae of Oriental species of Isonychia is also provided.

A new species of Cyclogomphus Selys, 1854 (Insecta: Odonata: Gomphidae) from the Western Ghats, India with comments on the status of Cyclogomphus vesiculosus Selys, 1873.

A new species of Cyclogomphus Selys, 1854 is described from the Western Ghats of India based on male and female specimens collected from Goa and Kerala states. The new species differs from all the known species of Cyclogomphus in the shape of the anal appendages as well as the thoracic and abdominal markings. We also provide here an updated key to the Cyclogomphus species and comment on the status of Cyclogomphus vesiculosus Selys, 1873.

A new species of Gomphidia Selys, 1854 (Insecta: Odonata: Anisoptera: Gomphidae) from the Western Ghats of India.

A new species of Gomphidia Selys, 1854 is described from southern Western Ghats, India based on a male specimen. The new species is very distinct from all the known species of Gomphidia from Indian subcontinent. Revised keys to the males and known females of Gomphidia of India and Sri Lanka are provided.

Review of the Cincticostella insolta (Allen, 1971) complex (Ephemeroptera: Ephemerellidae), with description of three new species from northern India and Nepal.

The Cincticostella insolta complex (Ephemeroptera: Ephemerellidae) is reviewed based on new and historic material, as well as new field observations. Three new species are described viz., C. richardi Martynov Palatov, sp. nov. and C. ranga Selvakumar Subramanian, sp. nov. from India, and C. sivaramakrishnani Martynov Palatov, sp. nov. from Nepal. A correction of the original description and some additional distinguishing features are given for C. braaschi Jacobus McCafferty, 2008. Cincticostella insolta (Allen, 1971), C. braaschi and C. bifurcata Xie, Jia, Chen, Jacobus Zhou, 2009 are recorded from the Indian subregion for the first time. Summaries of known distribution data and habitat preferences, as well as larval diagnoses, are presented for all seven species of the complex.