Phylogenetic divergences of the true bugs (Insecta: Hemiptera: Heteroptera), with emphasis on the aquatic lineages: the last piece of the aquatic insect jigsaw originated in the Late Permian/Early Triassic.

Heteroptera are among the most diverse hemimetabolous insects. Seven infraorders have been recognized within this suborder of Hemiptera. Apart from the well-established sister-group relationship between Cimicomorpha and Pentatomomorpha (= Terheteroptera), the two terminal lineages, the relationships among the other five infraorders are still controversial, of which three (Gerromorpha, Nepomorpha and Leptopodomorpha) are intimately connected to aquatic environments. However, the various and often conflicting available phylogeny hypotheses do not offer a clear background for a connection between diversification and palaeoenvironments. In this study, a molecular data set representing 79 taxa and 10 149 homologous sites is used to infer the phylogenetic relationships within Heteroptera. Bayesian inference, maximum-likelihood and maximum parsimony analyses were employed. The results of phylogenetic inferences largely confirm the widely accepted phylogenetic context. Estimation of the divergence time based on the phylogenetic results revealed that Gerromorpha, Nepomorpha and Leptopodomorpha originated successively during the period from the Late Permian to Early Triassic (269-246 Ma). This timescale is consistent with the origin and radiation time of various aquatic holometabolans. Our results indicate that the aquatic and semi-aquatic true bugs evolved under environmental conditions of high air temperature and humidity in an evolutionary scenario similar to that of the aquatic holometabolans.

Clavigeritae (Coleoptera, Staphylinidae, Pselaphinae) of the Arabian Peninsula with the description of a new species of Corynotopsis Jeannel, 1951 from Oman.

Corynotopsisomanicussp. nov. of the myrmecophilous supertribe Clavigeritae is described from Oman. The type series was collected at night and the ant host species remains unknown. Corynotopsisscotti Jeannel, 1951 is newly recorded for Yemen, and Commatocerusconcinnus Besuchet & Cuccodoro, 2011 for Oman. Lasius Fabricius, 1804 and Lepisiota Santschi, 1926 are, for the first time, determined as hosts of the latter species. The problematic taxonomic status of Corynotopsisscotti is discussed. A distribution map of all Clavigeritae known from the Arabian Peninsula is provided.

A new species of the genus Basidissus (Coleoptera: Anthribidae) from east Madagascar, with a key to species..

A new species, Basidissus bendai Trýzna & Baiat sp. nov. (Anthribidae: Anthribinae: Platyrhinini), from east Madagascar is described, female genitalia are studied and illustrated, and the mobility of the stylus of the toothed plate is discussed. Colour photographs as well as a key to Madagascan species of the genus Basidissus are provided.

Myrmicella, a new genus of Harpactorinae (Hemiptera: Heteroptera: Reduviidae) from Madagascar.

Myrmicella verticospinosa gen. et sp. nov., is described, based on four specimens (two males and two females) collected in south-west Madagascar (Zombitse-Vohibasia National Park and Isalo National Park). Three specimens have been collected by sifting the leaf litter, one female was collected using yellow pan traps. Genitalia of both sexes are described and illustrated.

New apterous Carventinae from Sri Lanka and Southern India (Hemiptera: Heteroptera: Aradidae).

Two new species of apterous Carventinae are described in this paper. One belongs to the endemic monotypic genus Singhalaptera Heiss 2001 from Sri Lanka (Singhalaptera secunda n.sp.), and the other to the monotypic genus Signocoris Hoberlandt 1958 from Madras (Signocoris nilgiricus n.sp.). All four known species of both genera are illustrated and compared.

A new species of the genus Apatenia (Coleoptera: Anthribidae) from Madagascar with notes on female genitalia, redescription of the female of Apatenia quadristigma Frieser and list of Madagascan species.

A new species, Apatenia stysi Trýzna & Banar sp. nov. (Anthribidae: Anthribinae: Corrhecerini), from north-west Mad-agascar is described and the female allotype of Apatenia quadristigma Frieser, 1981 is redescribed. Female genitalia of A. stysi sp. nov. are studied and illustrated.

Ambohitantelya yuripopovi gen. nov. et sp. nov., a new apterous Mezirinae from Madagascar (Hemiptera: Heteroptera: Aradidae) with unique metathoracic evaporatoria.

In addition to the ten genera of apterous Mezirinae recorded to date from Madagascar, a new genus Ambohitanlelya gen. nov. with uniquely formed metathoracic evaporatoria is erected for the A. yuripopovi sp. nov., which is described and illustrated. 

The Pselaphinae of Madagascar. II. Redescription of the genus Semiclaviger Wasmann, 1893 (Coleoptera: Staphylinidae: Pselaphinae: Clavigeritae) and synonymy of the subtribe Radamina Jeannel, 1954.

The enigmatic Madagascan genus Semiclaviger Wasmann is among the most morphologically distinct members of the obligately myrmecophilous pselaphine supertribe Clavigeritae. Here, the genus is redescribed, and the lectotype of the type species S. sikorae Wasmann is designated. We present a detailed study of the morphology of Semiclaviger, which supports its uniqueness among the Clavigeritae. The systematic position of the genus, and the validity of its subtribe Radamina, are discussed leading us to place Radamina in synonymy with Clavigerodina.

Pselaphinae of the Cameroon Volcanic Line. Part 2. Afroplectus cliveturneri, new name for Afroplectus turneri Hlavác & Banar, 2022 (Coleoptera, Staphylinidae) from the island São Tomé.

N/A.

Pselaphinae of the Cameroon Volcanic Line. Part 1. New Pselaphinae (Coleoptera, Staphylinidae) from the island São Tomé.

A new genus of Pselaphinae, Reichentomica gen. nov., and three new species, Reichentomica tasani sp. nov., Afroplectus turneri sp. nov., and Parasimus saotomicus sp. nov., are described for the first time from the island São Tomé. All three species were collected at the same locality close to Jardim Botânico do Bom Successo (00°16'31''N, 06°36'14''E) using flight intercept trap in the tropical forest at the elevation of 1324 m.

Description of a new species of the Anthaxia (Anthaxia) glabrifrons species-group from Morocco (Coleoptera: Buprestidae: Anthaxiini).

Anthaxia (Anthaxia) rejzeki sp. nov. belonging to the Anthaxia (Anthaxia) glabrifrons species-group from Morocco is described, illustrated and compared to other species of the group.

A contribution to the taxonomy of Anthaxia (Cratomerus) (Coleoptera: Buprestidae: Anthaxiini) of East Africa.

Anthaxia (Cratomerus) tankenethi sp. nov. from East Africa is described. New synonymy of A. (C.) gebhardti Obenberger, 1924 (= A. (C.) cytharaea Obenberger, 1931 syn. nov.) is proposed. Redescription of A. (C.) gebhardti is provided. Lectotypes for A. (C.) gebhardti Obenberger, 1924 and A. (C.) cytharaea Obenberger, 1931 are designated.

Two new species and new records of the genus Astenus from Madagascar (Coleoptera: Staphylinidae: Paederinae).

Two new species of the genus Astenus Dejean, 1833, A. ambohitantelyensis Janák Banar, sp. nov. and A. grebennikovi Janák Banar, sp. nov. from Madagascar are described and illustrated. New records of eleven additional species are presented.

Endogean and Cavernicolous Coleoptera of the Balkans. XXIII. A new species of Paramaurops J. Müller (Coleoptera: Staphylinidae: Pselaphinae) from Greece.

We describe and illustrate a new species of pselaphine beetles, Paramaurops zagoricus sp. n., from the Zagori region of Epirus, Greece. A list summarising the distribution of Amauropini known from Greece is also provided.

New species of Clavigeritae (Coleoptera: Staphylinidae: Pselaphinae) from Malawi and South Africa.

Three new species of Clavigeritae from subsaharan Africa, Zuluclavodes mulanjensis sp. nov., Zuluclavodes minor sp. nov. from southern Malawi and Articerodes jaloszynskii sp. nov. from South Africa, KwaZulu-Natal are described. Diagnosis is provided for both genera, and a key to species is given for the genus Zuluclavodes Hlavác, 2007. Ischyroceros mirus Reichensperger, 1915 is firstly recorded from Malawi, the illustration of its aedeagus is given. All described species are illustrated and their photos are presented.

A new species of Holophloeus Jordan (Coleoptera: Anthribidae) from eastern Madagascar with ecological notes on it and H. tuberosus (Fairmaire, 1897).

A new species, Holophloeus loebli Trýzna Banar sp. nov. (Anthribidae: Anthribinae: Discotenini), from east Madagascar is described. Male genitalia are studied and illustrated, and color photographs are provided. A comparison is provided with the other known Madagascan species of the genus, H. tuberosus (Fairmaire, 1897). Ecological notes, including color photographs of habitats, on H. loebli and H. tuberosus are provided.

First record of Systelloderes (Hemiptera: Heteroptera: Enicocephalidae) from Ecuador with a list of Ecuadorian Enicocephalomorpha.

The genus Systelloderes Blanchard, 1852 (Hemiptera: Heteroptera: Enicocephalomorpha: Enicocephalidae), has a nearly cosmopolitan distribution, with the greatest species richness being found in humid tropical and subtropical forests, but species are also present in humid microhabitats of temperate and arid zones (Wygodzinsky Schmidt 1991). In the Eastern Hemisphere species of Systelloderes occur in continental Africa (22 species, see Villiers 1969; 1976; Maldonado 1988; Banar 2008); Madagascar (Systelloders milloti Villiers, 1952); New Zealand (see Stys 1970, 2002) and New Caledonia (Systelloders loebli Stys Banar, 2007). Two species (S. capillicornis Bergroth, 1918 from Luzon and S. aetherius Bergroth, 1916 from Queensland) originally described as Systelloderes belong to the genus Henschiella Horváth, 1888 (P. Stys, unpublished data). As is frequently the case with Enicocephalomorpha, many species of Systelloderes remain to be described, especially from the Afrotropical, Neotropical and Oriental Regions. There are 13 described Systelloderes species from North and Central America (Wygodzinsky Schmidt 1991). To date, there are only six described species of Systelloderes from South America: two from Venezuela, and by a single species from Bolivia, Brazil, Chile and Peru each. There are very few additional records of Systelloderes in the New World tropics, including the works of Wolda (1975) and Ospina-Bautista (2018) on Colombia, Parker et al. (2012) on Peru, and Maestre et al. (2001) from Brazil.

Definition of Anthaxia (A.) hackeri species-group with description of a new species from China (Coleoptera: Buprestidae).

A new species Anthaxia (Anthaxia) tichyi sp. nov. from Yunnan, Southern China, is described, illustrated, including male genitalia and compared with the most similar species Anthaxia (A.) cupressi Bílý, 2005. The new species belongs to newly defined Anthaxia (Anthaxia) hackeri J. Frivaldszky, 1884 species-group.

A new Xenicocephalus species from Ecuador (Heteroptera, Enicocephalomorpha, Enicocephalidae).

Xenicocephalustomhenryi sp. n. (Insecta: Hemiptera: Heteroptera: Enicocephalomorpha: Enicocephalidae) is established for a single macropterous female from Ecuador. The enigmatic genus now includes three species known from only two Neotropical adults and an incomplete female specimen. The new species is described and illustrated, extensive comparative diagnoses for Xenicocephalus species are provided, and nomenclature, distribution, and biology of the genus are reviewed. The architecture of the raptorial forelegs of Xenicocephalus is unique among Enicocephalomorpha, and the genus is classified as subfamily incertae sedis.

A new Cervinotaptera species from northern Madagascar (Hemiptera, Heteroptera, Aradidae).

A new species, Cervinotapteratomhenryi, sp. n. (Hemiptera: Heteroptera: Aradidae: Mezirinae), from Montagne d'Ambre National Park in northern Madagascar is described and illustrated. The newly described species is compared with the only other known species, Cervinotapteraguilberti Heiss & Marchal, 2012.