A taxonomic revision of the Bothynus villiersi Endrdi, 1968 species group (Coleoptera: Scarabaeidae: Dynastinae).

The Bothynus villiersi species group (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) is revised and is composed of five South American species: Bothynus villiersi Endrdi, 1968; Bothynus horridus Endrdi, 1968; Bothynus alvarengai Martnez, 1983; Bothynus spinophallicus (Ratcliffe Cave, 2021) new combination; and Bothynus belemensis new species. Redescriptions, illustrations, new diagnostic characters, identification key, and a first known geographical distribution map for the group are presented.