Arthropods in modern resins reveal if amber accurately recorded forest arthropod communities.

Amber is an organic multicompound derivative from the polymerization of resin of diverse higher plants. Compared with other modes of fossil preservation, amber records the anatomy of and ecological interactions between ancient soft-bodied organisms with exceptional fidelity. However, it is currently suggested that ambers do not accurately record the composition of arthropod forest paleocommunities, due to crucial taphonomic biases. We evaluated the effects of taphonomic processes on arthropod entrapment by resin from the plant Hymenaea, one of the most important resin-producing trees and a producer of tropical Cenozoic ambers and Anthropocene (or subfossil) resins. We statistically compared natural entrapment by Hymenaea verrucosa tree resin with the ensemble of arthropods trapped by standardized entomological traps around the same tree species. Our results demonstrate that assemblages in resin are more similar to those from sticky traps than from malaise traps, providing an accurate representation of the arthropod fauna living in or near the resiniferous tree, but not of entire arthropod forest communities. Particularly, arthropod groups such as Lepidoptera, Collembola, and some Diptera are underrepresented in resins. However, resin assemblages differed slightly from sticky traps, perhaps because chemical compounds in the resins attract or repel specific insect groups. Ground-dwelling or flying arthropods that use the tree-trunk habitat for feeding or reproduction are also well represented in the resin assemblages, implying that fossil inclusions in amber can reveal fundamental information about biology of the past. These biases have implications for the paleoecological interpretation of the fossil record, principally of Cenozoic amber with angiosperm origin.

Molecular phylogeny of the spider family Sparassidae with focus on the genus Eusparassus and notes on the RTA-clade and 'Laterigradae'.

The phylogeny of the spider family Sparassidae is comprehensively investigated using four molecular markers (mitochondrial COI and 16S; nuclear H3 and 28S). Sparassidae was recovered as monophyletic and as most basal group within the RTA-clade. The higher-level clade Dionycha was not but monophyly of RTA-clade was supported. No affiliation of Sparassidae to other members of the 'Laterigradae' (Philodromidae, Selenopidae and Thomisidae) was observed, and the crab-like posture of this group assumed a result of convergent evolution. Only Philodromidae and Selenopidae were found members of a supported clade, but together with Salticidae and Corinnidae, while Thomisidae was nested within the higher Lycosoidea. Within Sparassidae monophyly of the subfamilies Heteropodinae sensu stricto, Palystinae and Deleninae was recovered. Sparianthinae was supported as the most basal clade within Sparassidae. Sparassinae and the genus Olios were found each to be polyphyletic. Eusparassinae was not recovered monophyletic, with the two original genera Eusparassus and Pseudomicrommata in separate clades and only the latter clustered with most other assumed Eusparassinae, here termed the "African clade". Further focus was on the monophyletic genus Eusparassus and its proposed species groups, of which the dufouri-, walckenaeri- and doriae-group were confirmed as monophyletic with the two latter groups more closely related. According to molecular clock analyses, the divergence time of Sparassidae and Eusparassus was estimated with 186 and 70 million years ago respectively.

Analysis of the soundscape in an intensive care unit based on the annotation of an audio recording.

The acoustic environments in hospitals, particularly in intensive care units (ICUs), are characterized by frequent high-level sound events which may negatively affect patient outcome. Many studies performed acoustic surveys, but the measurement protocol was not always reported in detail, and the scope of analysis was limited by the selected mode of sound level meters. Fewer studies systematically investigated the noise sources in ICUs by employing an observer in the patient room, which may potentially bias the measurement. In the current study, the soundscape of an ICU was evaluated where acoustic parameters were extracted from a ∼67-h audio recording, and a selected 24-h recording was annotated off-line for a source-specific analysis. The results showed that the patient-involved noise accounted for 31% of the acoustic energy and 11% of the predicted loudness peaks (PLPs). Excluding the patient-involved noise, the remaining acoustic energy was attributed to staff members (57%), alarms (30%), and the operational noise of life-supporting devices (13%). Furthermore, the contribution of each noise category to the PLPs was found to be more uneven: Staff (92%), alarms (6%), and device noise (2%). The current study suggests that most of the noise sources in ICUs may be associated with modifiable human factors.

Cebrennus Simon, 1880 (Araneae: Sparassidae): a revisionary up-date with the description of four new species and an updated identification key for all species.

The spider genus Cebrennus Simon, 1880 is revised again after thirteen years. Four new species are described: Cebrennus atlas spec. nov. from Morocco (female), C. flagellatus spec. nov. from Afghanistan (male), C. laurae spec. nov. from Canary Islands (male), and C. rechenbergi spec. nov. from Morocco (male and female). Cebrennus clercki (Audouin, 1826) comb. nov. is transferred from Philodromidae to Sparassidae and considered a nomen dubium. The holotype of C. aethiopicus Simon, 1880 is illustrated for the first time. Cebrennus tunetanus Simon, 1885 is re-described by illustrating its copulatory organs and some somatic characters, the internal duct system is shown for the first time supporting its placement in Cebrennus. An updated identification key for all species is provided. New records of Cebrennus species are listed: C. wagae (Simon, 1874) is recorded from Libya and Malta for the first time, the latter representing the first record for the entire genus from Europe. C. kochi (O. Pickard-Cambridge, 1872) is recorded from Syria, C. aethiopicus from Sudan for the first time. Records from the Canary Islands and from Afghanistan extend the known generic distribution range further to the West and East. Behavioural aspects (burrowing, escaping, mating) of C. rechenbergi and partly of C. villosus (Jézéquel & Junqua, 1966) are described. Photographs of this behaviour as well as of the habitus of several species are provided.

Six new species of the spider family Ochyroceratidae Fage 1912 (Arachnida: Araneae) from Southeast Asia.

Five new Althepus species and one new Psiloderces species of the family Ochyroceratidae are described from Southeast Asia: Althepus erectus spec. nov. (male) and A. nophaseudi spec. nov. (male, female) from Laos, A. flabellaris spec. nov. (male, female) from Thailand, A. reduncus spec. nov. (male) from Myanmar, A. spiralis spec. nov. (male) from Malaysia, and Psiloderces dicellocerus spec. nov. (male) from Indonesia. Primary types are deposited in the Senckenberg Research Institute in Frankfurt, Germany (SMF).

Four new species of pholcine spiders (Araneae: Pholcidae) from Southeast Asia.

Four new species belonging to four genera of the subfamily Pholcinae are reported from Southeast Asia: Belisana protumida spec. nov. (male, female), Khorata bayeri spec. nov. (male), Pholcus schawalleri spec. nov. (male), and Uthina khaosokensis spec. nov. (male).

Description of three new species of the spider genus Pseudopoda Jäger, 2000 (Araneae, Sparassidae) from China, Laos and Thailand, and the female of P.kavanaughi Zhang, Jäger & Liu, 2023.

With 252 species, Pseudopoda Jäger, 2000, is the largest genus in the family Sparassidae and is widely distributed in South (49 species in Bhutan, India, Nepal and Pakistan), East (158 species in China and Japan) and Southeast Asia (51 species in Indonesia, Laos, Myanmar, Thailand and Vietnam). Few species have been found in more than one region. In this paper, three new species of Pseudopoda are described from East and Southeast Asia. Among them, one from China: P.fengtongzhaiensis Jäger & Liu, sp. nov. (♀); one from Laos: P.baimai Jäger & Liu, sp. nov. (♀); and one from Thailand: P.inthanonensis Jäger & Liu, sp. nov. (♀). Additionally, the female of P.kavanaughi Zhang, Jäger & Liu, 2023 is described for the first time. Photos of the habitus and genitalia, as well as a distribution map of all four species, are provided.

Heteropodalebar sp. nov.: a new species from the highlands in Pahang State, Malaysia (Sparassidae, Heteropodinae) with a distinct sexual colour dimorphism.

Background: The genus Heteropoda Latreille, 1804, is ranked as the second within the family Sparassidae Bertkau, 1872. Up to now, sixteen species of this genus have been described from Malaysia. New information: A new species of this genus in the highlands of Pahang State, Malaysia is described under the name of H.lebar sp. nov.. Individuals of the new species live in primary forests on forest floor, active in the night on the leaf litter.

[Evidence-based practice in Bachelor of Nursing programmes in Austria, Germany, and Switzerland: A survey of general conditions, teaching content and methods]. / Evidenzbasierte Praxis in Bachelor-Pflegestudiengängen in Österreich, Deutschland und der Schweiz: ein Survey zu Rahmenbedingungen, Lehrinhalten und -methoden.

BACKGROUND/AIM: Evidence-based practice (EBP) provides an important basis for improving both the quality of care and patient safety. Formulating a research question, searching the literature, and critical appraisal are crucial to developing evidence-based practice. The aim of this survey was to provide an overview of how these topics are integrated into bachelor's degree programs in nursing in Austria, Germany, and the German-speaking part of Switzerland. We also aimed to show how teachers implement these subjects and how they experience and assess the implementation. METHOD: We conducted an exploratory cross-sectional study using an online survey sent out to program directors and teaching staff of all 58 bachelor's degree programs in nursing in Austria, Germany and the German-speaking part of Switzerland. For data collection, a questionnaire was developed containing items on general teaching conditions, contents, and methods of evidence-based nursing practice, as well as on the estimated thematic interest of students. The data were analysed descriptively. RESULTS: The program directors returned 24 questionnaires (41%). Of 75 questionnaires forwarded to the faculty, 17 (23%) were received from nine programs. On average, 5.6 teaching units (SD 2.6) are used for formulating a research question, 10 teaching units (SD 4.1) for literature review, and 11.3 teaching units (SD 6.9) for critical appraisal. Half of the teaching staff indicated that linkages between education and nursing care practice have been established. The traditional teaching method of frontal teaching is used predominantly. Student interest in topics was rated as moderate by most teachers. CONCLUSIONS: Topics on evidence-based practice are an integral part of bachelor's degree programs in nursing in German-speaking countries. An increase in teaching units, active learning methods and the growing interconnection between education and practice could improve the acquisition of competencies and attitudes of students regarding EBP and further advance its implementation in practice.

Anapidae (Arachnida: Araneae), a spider family newly recorded from Laos.

The spider family Anapidae is reported from Laos for the first time. One new species is described: Pseudanapis namkhan n. sp. from Nam Khan Valley, Luang Prabang Province. Another species is recorded for the first time from Champasak Province, Laos and the following new synonymy is proposed: Sinanapis thaleri Ono 2009 = Sinanapis crassitarsus Wunderlich & Song 1995 n. syn. Morphological descriptions, diagnoses and comparative illustrations are provided for the two species.

Nomenclatural and taxonomic problems related to the electronic publication of new nomina and nomenclatural acts in zoology, with brief comments on optical discs and on the situation in botany.

In zoological nomenclature, to be potentially valid, nomenclatural novelties (i.e., new nomina and nomenclatural acts) need first to be made available, that is, published in works qualifying as publications as defined by the International Code of zoological Nomenclature ("the Code"). In September 2012, the Code was amended in order to allow the recognition of works electronically published online after 2011 as publications available for the purpose of zoological nomenclature, provided they meet several conditions, notably a preregistration of the work in ZooBank. Despite these new Rules, several of the long-discussed problems concerning the electronic publication of new nomina and nomenclatural acts have not been resolved. The publication of this amendment provides an opportunity to discuss some of these in detail. It is important to note that: (1) all works published only online before 2012 are nomenclaturally unavailable; (2) printed copies of the PDFs of works which do not have their own ISSN or ISBN, and which are not obtainable free of charge or by purchase, do not qualify as publications but must be seen as facsimiles of unavailable works and are unable to provide nomenclatural availability to any nomenclatural novelties they may contain; (3) prepublications online of later released online publications are unavailable, i.e., they do not advance the date of publication; (4) the publication dates of works for which online prepublications had been released are not those of these prepublications and it is critical that the real release date of such works appear on the actual final electronic publication, but this is not currently the case in electronic periodicals that distribute such online prepublications and which still indicate on their websites and PDFs the date of release of prepublication as that of publication of the work; (5) supplementary online materials and subsequent formal corrections of either paper or electronic publications distributed only online are nomenclaturally unavailable; (6) nomenclatural information provided on online websites that do not have a fixed content and format, with ISSN or ISBN, is unavailable. We give precise examples of many of these nomenclatural problems. Several of them, when they arise, are due to the fact that the availability of nomenclatural novelties now depends on information that will have to be sought not from the work itself but from extrinsic evidence. As shown by several examples discussed here, an electronic document can be modified while keeping the same DOI and publication date, which is not compatible with the requirements of zoological nomenclature. Therefore, another system of registration of electronic documents as permanent and inalterable will have to be devised. ZooBank also clearly needs to be improved in several respects. Mention in a work of its registration number (LSID) in ZooBank would seem to be possible only if this registration has occurred previously, but some works that have purportedly been registered in ZooBank are in fact missing on this web application. In conclusion, we offer recommendations to authors, referees, editors, publishers, libraries and the International Commission on Zoological Nomenclature, in the hope that such problems can be limited along with the potential chaos in zoological nomenclature that could result, if careful attention is not paid to the problems we highlight here, from a somewhat misplaced, and perhaps now widespread, understanding that electronic publication of nomenclatural novelties is now allowed and straightforward. We suggest that, as long as the problematic points linked to the new amendment and to electronic publication as a whole are not resolved, nomenclatural novelties continue to be published in paper-printed journals that have so far shown editorial competence regarding taxonomy and nomenclature, which is not the case of several recent electronic-only published journals.

Geometric regularity in webs of non-orb-weaving spiders.

Geometric regularity of spider webs has been intensively studied in orb-weaving spiders, although it is not exclusive of orb weavers. Here, we document the geometrically regular, repetitive elements in the webs of the non-orb-weaving groups Leptonetidae and Telemidae for the first time. Similar to orb weavers, we found areas with regularly spaced parallel lines in the webs of Calileptoneta helferi, Sulcia sp., and cf. Pinelema sp. Furthermore, we provide a detailed account of the regular webs of Ochyrocera (Ochyroceratidae). The sections of the web with regularly disposed parallel lines are built as U-shaped modules reminiscent of orb webs. It has been suggested that the regularly spaced parallel lines in the webs of Ochyroceratidae and Psilodercidae may be produced in a single sweep of their posterior lateral spinnerets, which have regularly spaced aciniform gland spigots, perhaps involving expansion of the spinnerets. To test this hypothesis, we compared the spacing between parallel lines with the spacing between spigots, searched for expansible membranes in the spinnerets, and examined the junctions of regularly spaced lines. The distance between parallel lines was 10-20 times the distance between spigots, and we found no expansible membranes, and the intersection of parallel lines are cemented, which opposes the single sweep hypothesis. Furthermore, we found cues of viscid silk in the parallel lines of the psilodercid Althepus and broadened piriform gland spigots that may be responsible of its production. Finally, we evaluated the presence or absence of geometrically regular web elements across the spider tree of life. We found reports of regular webs in 31 spider families, including 20 families that are not orb weavers and hypothesize that the two basic aspects of regularity (parallel lines spaced at regular intervals, and radial lines spaced at regular angles) probably appeared many times in the evolution of spiders.

Computed tomography recovers data from historical amber: an example from huntsman spiders.

Computed tomography (CT) methods were applied to a problematic fossil spider (Arachnida: Araneae) from the historical Berendt collection of Eocene (ca. 44-49 Ma) Baltic amber. The original specimens of Ocypete crassipes Koch and Berendt 1854 are in dark, oxidised amber and the published descriptions lack detail. Despite this, they were subsequently assigned to the living Pantropical genus Heteropoda Latreille, 1804 and are ostensibly the oldest records of huntsman spiders (Sparassidae) in general. Given their normally large size, and presumptive ability to free themselves more easily from resin, it would be surprising to find a sparassid in amber and traditional (optical) methods of study would likely have left O. crassipes as an equivocal record--probably a nomen dubium. However, phase contrast enhanced X-ray CT revealed exquisite morphological detail and thus 'saved' this historical name by revealing characters which confirm that it's a bona fide member both of Sparassidae and the subfamily Eusparassinae. We demonstrate here that CT studies facilitate taxonomic equivalence even between recent spiders and unpromising fossils described in older monographs. In our case, fine structural details such as eye arrangement, cheliceral dentition, and leg characters like a trilobate membrane, spination and claws, allow a precise referral of this fossil to an extant genus as Eusparassus crassipes (Koch and Berendt 1854) comb. nov.

Two new enigmatic genera of huntsman spiders from Madagascar (Araneae: Sparassidae).

Two new monotypic genera of the family Sparassidae are described from Madagascar: Martensikara gen. nov. with M. jocheni spec. nov. (female; from Toliara Province), belonging to Heteropodinae Thorell, 1873, and Deelemanikara gen. nov. with D. christae spec. nov. (female; from Toamasina and Antananarivo Provinces) of uncertain systematic affiliation. Both genera show unique combinations of characters that do not occur in any of the known genera. Somatic as well as genital characters of both genera are discussed and compared with those of other taxa of Sparassidae.

Eight legs, two wings, one life for scientific research-celebrating the 80th birthday of Prof. Dr Jochen Martens.

The scientific life of Prof. Dr Jochen Martens (Germany: Mainz) is illuminated on occasion of his 80th birthday. Facts and impressions are given as well as lists of his publications (329), taxa he has described (2 families, 29 genera, 296 species) and that have been dedicated to him (11 genera, 219 species, 1 subspecies). Jochen Martens is a renowned specialist for birds (Aves) and for harvestmen (Opiliones). So far, he travelled to 27 countries in 80 journeys with Nepal and the Himalayas as one geographic focus.

Coelotesvignai Brignoli, 1978 (Araneae: Agelenidae) from Turkey: first description of male and annotations on terminology of copulatory organs.

BACKGROUND: The agelenid spider species Coelotesvignai Brignoli, 1978 was described, based on female specimens from Turkey. NEW INFORMATION: The unknown male is here described, based on specimens from the type locality: Bolu, Abant Mountains, Turkey. The variation of the female copulatory organs is illustrated. The relationships of the species with its putative closest congeners are discussed. The discrepancy between the morphological terminology used in the Coelotinae and Ageleninae is discussed and some suggestions how to unify them are proposed.

Preface.

To process a Festschrift is an honour and a unique opportunity to show respect to someone who made important contributions over many years in a special field of scientific research. In the case of Jochen Martens the contribution is to not only in one field, but in several: harvestmen, birds, high mountains, taxonomy, systematics, evolution etc.

Pholcid spiders (Araneae: Pholcidae) from Southeast Asia, with descriptions of five new species.

Five new pholcid species belonging to Holocneminus Berland, 1942, Khorata Huber, 2005 and Pholcus Walckenaer, 1805 are newly described from Southeast Asia: Holocneminus samanggi Lan Li sp. nov. (Indonesia, male and female), Khorata kep Lan, Jäger Li sp. nov. (Cambodia, male), Khorata musee Lan Li sp. nov. (Thailand, male and female), Pholcus bat Lan Li sp. nov. (China, male and female), and Pholcus phnombak Lan, Jäger Li sp. nov. (Cambodia, male and female). Species from the genera Khorata and Pholcus are reported from Cambodia for the first time.

Twenty years, eight legs, one concept: describing spider biodiversity in Zootaxa (Arachnida: Araneae).

Zootaxa published more than a thousand papers on Araneae from 2002 to the present, including descriptions of 3,833 new spider species and 177 new genera. Here we summarise the key contributions of Zootaxa to our current knowledge of global spider diversity. We provide a historical account of the researchers that have actively participated as editors, and recognize the more than 1,000 reviewers without whom none of this would have been possible. We conduct a simple analysis of the contributions by authors and geographic region, which allows us to uncover some of the underlying trends in current spider taxonomy. In addition, we examine some of the milestones in twenty years of spider systematic research in Zootaxa. Finally, we discuss future prospects of spider taxonomy and the role that Zootaxa and its younger sister journal Megataxa will play in it. We would like to dedicate this contribution to the memory of Norman I. Platnick, a crucial figure in the advancement of spider systematics.

The spider genus Olios Walckenaer, 1837 (Araneae: Sparassidae)-Part 1: species groups, diagnoses, identification keys, distribution maps and revision of the argelasius-, coenobitus- and auricomis-groups.

The genus Olios Walckenaer, 1837 is revised, a generic diagnosis is given and an identification key to eight species groups is provided. Olios in its revised sense includes 87 species and is distributed in Africa, southern Europe and Asia. Three species groups are revised in this first part, an identification key to species for each group is provided, five new species are described and all included species are illustrated. The Olios argelasius-group includes O. argelasius Walckenaer, 1806, O. canariensis (Lucas, 1838), O. pictus (Simon, 1885), O. fasciculatus Simon, 1880 and O. kunzi spec. nov. (male, female; Namibia, Zambia, South Africa); it is distributed in the Mediterranean region, northern Africa including Canary Islands, in the Middle East, South Sudan, East Africa, and southern Africa. The Olios coenobitus-group includes O. angolensis spec. nov. (male; Angola), O. coenobitus Fage, 1926, O. denticulus spec. nov. (male; Java), O. erraticus Fage, 1926, O. gambiensis spec. nov. (male, female; Gambia), O. milleti (Pocock, 1901b), O. mordax (O. Pickard-Cambridge, 1899) and O. pusillus Simon, 1880; it is distributed in Africa (Gambia, Angola, Tanzania, Madagascar) and Asia (India, Sri Lanka, Indonesia: Java). The Olios auricomis-group includes only O. auricomis (Simon, 1880), distributed in Africa south of 10°N. Other species groups are introduced briefly and will be revised in forthcoming revisions. The Olios correvoni-group includes currently O. claviger (Pocock, 1901a), O. correvoni Lessert, 1921, O. correvoni choupangensis Lessert, 1936, O. darlingi (Pocock, 1901a), O. faesi Lessert, 1933, O. freyi Lessert, 1929, O. kassenjicola Strand, 1916b, O. kruegeri (Simon, 1897a), O. quadrispilotus (Simon, 1880) comb. nov., O. lucieni comb. nov. nom. nov., O. sjostedti Lessert, 1921 and O. triarmatus Lessert, 1936; it is distributed in Africa (Zimbabwe, Tanzania incl. Zanzibar, Angola, Congo, Central Africa, South Africa, Botswana; O. darlingi was recorded from Zimbabwe and Botswana and not from South Africa). The Olios rossettii-group includes: O. baulnyi (Simon, 1874), O. bhattacharjeei (Saha Raychaudhuri, 2007), O. brachycephalus Lawrence, 1938, O. floweri Lessert, 1921, O. jaldaparaensis Saha Raychaudhuri, 2007, O. japonicus Jäger Ono, 2000, O. kolosvaryi (Caporiacco, 1947b) comb. nov., O. longipes (Simon, 1884b), O. lutescens (Thorell, 1894), O. mahabangkawitus Barrion Litsinger, 1995, O. obesulus (Pocock, 1901b), O. rossettii (Leardi, 1901), O. rotundiceps (Pocock, 1901b), O. sericeus (Kroneberg, 1875), O. sherwoodi Lessert, 1929, O. suavis (O. Pickard-Cambridge, 1876), O. tarandus (Simon, 1897d), O. tener (Thorell, 1891) and O. tiantongensis (Zhang Kim, 1996); it is distributed in the Mediterranean region, in Africa (especially eastern half) and Asia (Middle East and Central Asia to Japan, Philippines and Java). The Olios nentwigi-group includes O. diao Jäger, 2012, O. digitatus Sun, Li Zhang, 2011, O. jaenicke Jäger, 2012, O. muang Jäger, 2012, O. nanningensis (Hu Ru, 1988), O. nentwigi spec. nov. (male, female; Indonesia: Krakatau), O. perezi Barrion Litsinger, 1995, O. scalptor Jäger Ono, 2001 and O. suung Jäger, 2012; it is distributed in Asia (Thailand, Laos, Vietnam, Cambodia, China, Taiwan, Indonesia, Philippines), Papua New Guinea and Mariana Islands. Olios diao is newly recorded from Cambodia and Champasak Province in Laos. The Olios stimulator-group includes O. admiratus (Pocock, 1901b), O. hampsoni (Pocock, 1901b), O. lamarcki (Latreille, 1806) and O. stimulator Simon, 1897c; it is distributed in Africa (Madagascar, Seychelles), Middle East and South Asia (United Arab Emirates, Iraq, Afghanistan, Pakistan, India, Maldives, Sri Lanka). The Olios hirtus-group includes O. bungarensis Strand, 1913b, O. debalae (Biswas Roy, 2005), O. ferox (Thorell, 1892), O. hirtus (Karsch, 1879a), O. igraya (Barrion Litsinger, 1995) comb. nov., O. menghaiensis (Wang Zhang, 1990), O. nigrifrons (Simon, 1897b), O. punctipes Simon, 1884a, O. punctipes sordidatus (Thorell, 1895), O. pyrozonis (Pocock, 1901b), O. sungaya (Barrion Litsinger, 1995) comb. nov., O. taprobanicus Strand, 1913b and O. tikaderi Kundu et al., 1999; it is distributed in South, East and Southeast Asia (Sri Lanka, India, Nepal, Bangladesh, Myanmar, China, Laos, Thailand, Cambodia, Vietnam, Malaysia, Indonesia, Philippines). Nineteen synonyms are recognised: Nisueta Simon, 1880, Nonianus Simon, 1885, both = Olios syn. nov.; O. spenceri Pocock, 1896, O. werneri (Simon, 1906a), O. albertius Strand, 1913a, O. banananus Strand, 1916a, O. aristophanei Lessert, 1936, all = O. fasciculatus; O. subpusillus Strand, 1907c = O. pusillus; O. schonlandi (Pocock, 1900b), O. rufilatus Pocock, 1900c, O. chiracanthiformis Strand, 1906, O. ituricus Strand, 1913a, O. isongonis Strand, 1915, O. flavescens Caporiacco, 1941 comb. nov., O. pacifer Lessert, 1921, all = O. auricomis; Olios sanguinifrons (Simon, 1906b) = O. rossettii Leardi, 1901; O. phipsoni (Pocock, 1899), Sparassus iranii (Pocock, 1901b), both = O. stimulator; O. fuligineus (Pocock, 1901b) = O. hampsoni. Nine species are transferred to Olios: O. gaujoni (Simon, 1897b) comb. nov., O. pictus comb. nov., O. unilateralis (Strand, 1908b) comb. nov. (all three from Nonianus), O. affinis (Strand, 1906) comb. nov., O. flavescens Caporiacco, 1941 comb. nov., O. quadrispilotus comb. nov., O. similis (Berland, 1922) comb. nov. (all four from Nisueta), O. sungaya (Barrion Litsinger, 1995) comb. nov., O. igraya (Barrion Litsinger, 1995) comb. nov. (both from Isopeda L. Koch 1875). Olios lucieni nom. nov. comb. nov. is proposed for Nisueta similis Berland, 1922, which becomes a secondary homonym. The male of O. quadrispilotus comb. nov. is described for the first time. Sixteen species are currently without affiliation to one of the eight species groups: O. acolastus (Thorell, 1890), O. alluaudi Simon, 1887a, O. batesi (Pocock, 1900c), O. bhavnagarensis Sethi Tikader, 1988, O. croseiceps (Pocock, 1898b), O. durlaviae Biswas Raychaudhuri, 2005, O. gentilis (Karsch, 1879b), O. gravelyi Sethi Tikader, 1988, O. greeni (Pocock, 1901b), O. inaequipes (Simon 1890), O. punjabensis Dyal, 1935, O. ruwenzoricus Strand, 1913a, O. senilis Simon, 1880, O. somalicus Caporiacco, 1940, O. wroughtoni (Simon, 1897c) and O. zulu Simon, 1880. Five of these species are illustrated in order to allow identification of the opposite (male) sex and to settle their systematic placement. Thirty-seven species are considered nomina dubia, mostly because they were described from immatures, three of them are illustrated: O. abnormis (Blackwall, 1866), O. affinis (Strand, 1906) comb. nov., O. africanus (Karsch, 1878), O. amanensis Strand, 1907a, O. annandalei (Simon, 1901), O. bivittatus Roewer, 1951, O. ceylonicus (Leardi, 1902), O. conspersipes (Thorell, 1899), Palystes derasus (C.L. Koch, 1845) comb. nov., O. detritus (C.L. Koch, 1845), O. digitalis Eydoux Souleyet, 1842, O. exterritorialis Strand, 1907b, O. flavovittatus (Caporiacco, 1935), O. fugax (O. Pickard-Cambridge, 1885), O. guineibius Strand, 1911c, O. guttipes (Simon, 1897a), O. kiranae Sethi Tikader, 1988, O. longespinus Caporiacco, 1947b, O. maculinotatus Strand, 1909, O. morbillosus (MacLeay, 1827), O. occidentalis (Karsch, 1879b), O. ornatus (Thorell, 1877), O. pagurus Walckenaer, 1837, O. patagiatus (Simon, 1897b), O. praecinctus (L. Koch, 1865), O. provocator Walckenaer, 1837, O. quesitio Moradmand, 2013, O. quinquelineatus Taczanowski, 1872, O. sexpunctatus Caporiacco, 1947a, Heteropoda similaris (Rainbow, 1898) comb. rev., O. socotranus (Pocock, 1903), O. striatus (Blackwall, 1867), O. timidus (O. Pickard-Cambridge, 1885), Remmius variatus (Thorell, 1899) comb. nov., O. vittifemur Strand, 1916b, O. wolfi Strand, 1911a and O. zebra (Thorell, 1881). Eighty-nine species are misplaced in Olios but cannot be affiliated to any of the known genera. They belong to the subfamilies Deleninae Hogg, 1903, Sparassinae Bertkau, 1872 and Palystinae Simon, 1897a, nineteen of them are illustrated: O. acostae Schenkel, 1953, O. actaeon (Pocock, 1898c), O. artemis Hogg, 1915, O. atomarius Simon, 1880, O. attractus Petrunkevitch, 1911, O. auranticus Mello-Leitão, 1918, O. benitensis (Pocock, 1900c), O. berlandi Roewer, 1951, O. biarmatus Lessert, 1925, O. canalae Berland, 1924, O. caprinus Mello-Leitão, 1918, O. chelifer Lawrence, 1937, O. chubbi Lessert, 1923, O. clarus (Keyserling, 1880), O. coccineiventris (Simon, 1880), O. corallinus Schmidt, 1971, O. crassus Banks, 1909, O. debilipes Mello-Leitão, 1945, O. discolorichelis Caporiacco, 1947a, O. erroneus O. Pickard-Cambridge, 1890, O. extensus Berland, 1924, O. fasciiventris Simon, 1880 , O. feldmanni Strand, 1915, O. fimbriatus Chrysanthus, 1965, O. flavens Nicolet, 1849, O. fonticola (Pocock, 1902), O. formosus Banks, 1929, O. francoisi (Simon, 1898a), O. fulvithorax Berland, 1924, O. galapagoensis Banks, 1902, O. gaujoni (Simon, 1897b) comb. nov., O. giganteus Keyserling, 1884, O. hoplites Caporiacco, 1941, O. humboldtianus Berland, 1924, O. insignifer Chrysanthus, 1965, O. insulanus (Thorell, 1881), O. keyserlingi (Simon, 1880), O. lacticolor Lawrence, 1952, O. lepidus Vellard, 1924, O. longipedatus Roewer, 1951, O. machadoi Lawrence, 1952, O. macroepigynus Soares, 1944, O. maculatus Blackwall, 1862, O. marshalli (Pocock, 1898a), O. mathani (Simon, 1880), O. minensis Mello-Leitão, 1917, O. monticola Berland, 1924, O. mutabilis Mello-Leitão, 1917, O. mygalinus Doleschall, 1857, O. mygalinus cinctipes Merian, 1911, O. mygalinus nirgripalpis Merian, 1911, O. neocaledonicus Berland, 1924, O. nigristernis (Simon, 1880), O. nigriventris Taczanowski, 1872, O. oberzelleri Kritscher, 1966, O. obscurus (Keyserling, 1880), O. obtusus F.O. Pickard-Cambridge, 1900, O. orchiticus Mello-Leitão, 1930, O. oubatchensis Berland, 1924, O. paraensis (Keyserling, 1880), O. pellucidus (Keyserling, 1880), O. peruvianus Roewer, 1951, O. pictitarsis Simon, 1880, O. plumipes Mello-Leitão, 1937, O. princeps Hogg, 1914, O. pulchripes (Thorell, 1899), O. puniceus (Simon, 1880), O. roeweri Caporiacco, 1955a, O. rubripes Taczanowski, 1872, O. rubriventris (Thorell, 1881), O. rufus Keyserling, 1880, O. sanctivincenti (Simon, 1898b), O. similis (O. Pickard-Cambridge, 1890), O. simoni (O. Pickard-Cambridge, 1890), O. skwarrae Roewer, 1933, O. spinipalpis (Pocock, 1901a), O. stictopus (Pocock, 1898a), O. strandi Kolosváry, 1934, O. subadultus Mello-Leitão, 1930, O. sulphuratus (Thorell, 1899), O. sylvaticus (Blackwall, 1862), O. tamerlani Roewer, 1951, O. tigrinus (Keyserling, 1880), O. trifurcatus (Pocock, 1900c), O. trinitatis Strand, 1916a, O. velox (Simon, 1880), O. ventrosus Nicolet, 1849, O. vitiosus Vellard, 1924 and O. yucatanus Chamberlin, 1925. Seventeen taxa are transferred from Olios to other genera within Sparassidae, eight of them are illustrated: Adcatomus luteus (Keyserling, 1880) comb. nov., Eusparassus flavidus (O. Pickard-Cambridge, 1885) comb. nov., Palystes derasus (C.L. Koch, 1845) comb. nov., Heteropoda similaris (Rainbow, 1898) comb. rev., Remmius variatus (Thorell, 1899) comb. nov., Nolavia audax (Banks, 1909) comb. nov., Nolavia antiguensis (Keyserling, 1880) comb. nov., Nolavia antiguensis columbiensis (Schmidt, 1971) comb. nov., Nolavia fuhrmanni (Strand, 1914) comb. nov., Nolavia helva (Keyserling, 1880) comb. nov., Nolavia stylifer (F.O. Pickard-Cambridge, 1900) comb. nov., Nolavia valenciae (Strand, 1916a) comb. nov., Nungara cayana (Taczanowski, 1872) comb. nov., Polybetes bombilius (F.O. Pickard-Cambridge, 1899) comb. nov., Polybetes fasciatus (Keyserling, 1880) comb. nov., Polybetes hyeroglyphicus (Mello-Leitão, 1918) comb. nov. and Prychia paalonga (Barrion Litsinger, 1995) comb. nov. One species is transferred from Olios to the family Clubionidae Wagner, 1887: Clubiona paenuliformis (Strand, 1916a) comb. nov.