Heterogeneous contributions can jeopardize cooperation in the public goods game.

When studying social dilemma games, a crucial question arises regarding the impact of general heterogeneity on cooperation, which has been shown to have positive effects in numerous studies. Here, we demonstrate that heterogeneity in the contribution value for the focal public goods game can jeopardize cooperation. We show that there is an optimal contribution value in the homogeneous case that most benefits cooperation depending on the lattice. In a heterogeneous scenario, where strategy and contribution coevolve, cooperators making contributions higher than the optimal value end up harming those who contribute less. This effect is notably detrimental to cooperation in the square lattice with von Neumann neighborhood, while it can have no impact in other lattices. Furthermore, in parameter regions where a higher-contributing cooperator cannot normally survive alone, the exploitation of lower-value contribution cooperators allows their survival, resembling a parasitic behavior. To obtain these results, we examined the effect of various distributions for the contribution values in the initial condition and we conducted Monte Carlo simulations.

Symbiotic behaviour in the public goods game with altruistic punishment.

Finding ways to overcome the temptation to exploit one another is still a challenge in behavioural sciences. In the framework of evolutionary game theory, punishing strategies are frequently used to promote cooperation in competitive environments. Here, we introduce altruistic punishers in the spatial public goods game. This strategy acts as a cooperator in the absence of defectors, otherwise it will punish all defectors in their vicinity while bearing a cost to do so. We observe three distinct behaviours in our model: i) in the absence of punishers, cooperators (who don't punish defectors) are driven to extinction by defectors for most parameter values; ii) clusters of punishers thrive by sharing the punishment costs when these are low; iii) for higher punishment costs, punishers, when alone, are subject to exploitation but in the presence of cooperators can form a symbiotic spatial structure that benefits both. This last observation is our main finding since neither cooperation nor punishment alone can survive the defector strategy in this parameter region and the specificity of the symbiotic spatial configuration shows that lattice topology plays a central role in sustaining cooperation. Results were obtained by means of Monte Carlo simulations on a square lattice and subsequently confirmed by a pairwise comparison of different strategies' payoffs in diverse group compositions, leading to a phase diagram of the possible states.