Early North African Acheulean techno-economic systems at Thomas Quarry I - L1 (Casablanca, Morocco).

North Africa is a key area for understanding cultural processes that led to the Acheulean pan - African emergence and expressions and the related hominin population dynamics. Unfortunately, little is known about the early Acheulean in this vast area of the African continent due to the scarceness of archaeological sites in stratigraphic context with reliable chronometric data, human remains, and technological analyses of the lithic industries. Here, we present the first comprehensive techno-economic analysis of the early Acheulean assemblage from Thomas Quarry I - Unit L1 (ThI-L1, Casablanca, Morocco), which is the earliest Acheulean site of North Africa, unambiguously dated to 1.3 Ma. Fieldwork has unearthed faunal remains and a lithic collection containing over 3800 artefacts, which represents one of the largest series for the early African Acheulean. The assemblage is mainly composed of quartzites and to a lesser extent of silicites, both abundantly available near the site. Previously published results of the silicite study revealed two different productions for the extraction of small flakes and of bladelet-like flakes. In this work, we analyse the techno-economic systems of the quartzite assemblage. Two distinct quartzite productions co-occur, one devoted to the manufacture of Large Cutting Tools (LCTs), the other focused on the extraction of small-medium sized flakes. LCTs were usually produced from large cobbles, less often from large flakes detached mainly using the entame core method. The main technical objective was to obtain large pointed tools and, more rarely, large tools with a transversal cutting edge. Results support the existence of a strong synergy between conceptual and operational schemes regulated by the ability to anticipate the final tool morphology and to apply standardized shaping procedures to manufacture recurrent morphotypes. Quartzite small-medium sized flakes were produced by a diversity of flaking methods adapted to the cobble blank morphologies and were not retouched. The results allow to assess that the earliest technical expression of the Acheulean in North Africa is characterised by a high diversification of the stone knapping outcomes, the complexity of the mental templates, and the flexible structure of the operational schemes.

First high resolution chronostratigraphy for the early North African Acheulean at Casablanca (Morocco).

The onset of the Acheulean, marked by the emergence of large cutting tools (LCTs), is considered a major technological advance in the Early Stone Age and a key turning point in human evolution. The Acheulean originated in East Africa at ~ 1.8-1.6 Ma and is reported in South Africa between ~ 1.6 and > 1.0 Ma. The timing of its appearance and development in North Africa have been poorly known due to the near-absence of well-dated sites in reliable contexts. The ~ 1 Ma stone artefacts of Tighennif (Algeria) and Thomas Quarry I-Unit L (ThI-L) at Casablanca (Morocco) are thus far regarded as documenting the oldest Acheulean in North Africa but whatever the precision of their stratigraphical position, both deserve a better chronology. Here we provide a chronology for ThI-L, based on new magnetostratigraphic and geochemical data. Added to the existing lithostratigraphy of the Casablanca sequence, these results provide the first robust chronostratigraphic framework for the early North African Acheulean and firmly establish its emergence in this part of the continent back at least to ~ 1.3 Ma.

An ecological niche shift for Neanderthal populations in Western Europe 70,000 years ago.

Middle Paleolithic Neanderthal populations occupied Eurasia for at least 250,000 years prior to the arrival of anatomically modern humans. While a considerable body of archaeological research has focused on Neanderthal material culture and subsistence strategies, little attention has been paid to the relationship between regionally specific cultural trajectories and their associated existing fundamental ecological niches, nor to how the latter varied across periods of climatic variability. We examine the Middle Paleolithic archaeological record of a naturally constrained region of Western Europe between 82,000 and 60,000 years ago using ecological niche modeling methods. Evaluations of ecological niche estimations, in both geographic and environmental dimensions, indicate that 70,000 years ago the range of suitable habitats exploited by these Neanderthal populations contracted and shifted. These ecological niche dynamics are the result of groups continuing to occupy habitual territories that were characterized by new environmental conditions during Marine Isotope Stage 4. The development of original cultural adaptations permitted this territorial stability.

The origin of early Acheulean expansion in Europe 700 ka ago: new findings at Notarchirico (Italy).

Notarchirico (Southern Italy) has yielded the earliest evidence of Acheulean settlement in Italy and four older occupation levels have recently been unearthed, including one with bifaces, extending the roots of the Acheulean in Italy even further back in time. New 40Ar/39Ar on tephras and ESR dates on bleached quartz securely and accurately place these occupations between 695 and 670 ka (MIS 17), penecontemporaneous with the Moulin-Quignon and la Noira sites (France). These new data demonstrate a very rapid expansion of shared traditions over Western Europe during a period of highly variable climatic conditions, including interglacial and glacial episodes, between 670 and 650 (i.e., MIS17/MIS16 transition). The diversity of tools and activities observed in these three sites shows that Western Europe was populated by adaptable hominins during this time. These conclusions question the existence of refuge areas during intense glacial stages and raise questions concerning understudied migration pathways, such as the Sicilian route.

Dedicated core-on-anvil production of bladelet-like flakes in the Acheulean at Thomas Quarry I - L1 (Casablanca, Morocco).

The ability to produce large cutting tools (LCTs) is considered as the technological marker of the Acheulean and the indicator of a greater technological complexity compared to the previous Oldowan. Although Acheulean techno-complexes are also composed of a concurrent core-and-flake technology, the iconic handaxes have attracted more attention than any other lithic component. Consequently, little is known of the small and medium-sized flake productions (small flaking), especially starting from 1 Ma, when handaxe and cleaver manufacture becomes intensive and widespread across Africa, including the Atlantic coastal regions of Morocco. Research at Thomas Quarry I yielded a rich early Acheulean lithic assemblage, mainly composed of quartzite LCTs and small flaking, together with a small-sized flint production. Here, we report a particular aspect of this flint assemblage, i.e. a flint bladelet-like flake production. This process represents a discrete technical behaviour among those related to small flaking both in quartzite and flint: pebbles were flaked using the bipolar-on-anvil technique repeatedly employing a specific method to produce bladelet-like flakes. This production represents the oldest dated occurrence of bladelet-like technology in Africa and reveals technical competencies hitherto unknown for these periods, providing further elements for the techno-economic diversification of the African Acheulean.

Earliest African evidence of carcass processing and consumption in cave at 700 ka, Casablanca, Morocco.

To date, in Africa, evidence for animal processing and consumption in caves routinely used as living spaces is only documented in the late Middle Pleistocene of the North and South of the continent and postdates the Middle Pleistocene in East Africa. Here we report the earliest evidence in a North-African cave (Grotte des Rhinocéros at Casablanca, Morocco) of cut, percussion and human gnawing marks on faunal remains directly associated with lithic knapping activities in the same space and in a well-documented stratified context. Ages for this Acheulean site are provided by the dating of herbivorous teeth to 690-720 ka and 520-550 ka (lower and upper sets) by combined Electron Spin Resonance (ESR) and U-series techniques. Traces of butchery on gazelle, alcelaphin, and zebra bones demonstrate that hominins had primary access to herbivore carcasses. Hominins brought and consumed meat in the cave, as documented by herbivore bones bearing human tooth marks concentrated in a circumscribed area of the excavation. In Africa, this site provides the earliest evidence for in situ carcass processing and meat-eating in cave, directly associated with lithic production and demonstrates the recurrent use by early Middle Pleistocene hominins of a North African cave site 400 000 years before that by Homo sapiens at Jebel Irhoud (Morocco).

The age of the hominin fossils from Jebel Irhoud, Morocco, and the origins of the Middle Stone Age.

The timing and location of the emergence of our species and of associated behavioural changes are crucial for our understanding of human evolution. The earliest fossil attributed to a modern form of Homo sapiens comes from eastern Africa and is approximately 195 thousand years old, therefore the emergence of modern human biology is commonly placed at around 200 thousand years ago. The earliest Middle Stone Age assemblages come from eastern and southern Africa but date much earlier. Here we report the ages, determined by thermoluminescence dating, of fire-heated flint artefacts obtained from new excavations at the Middle Stone Age site of Jebel Irhoud, Morocco, which are directly associated with newly discovered remains of H. sapiens. A weighted average age places these Middle Stone Age artefacts and fossils at 315 ± 34 thousand years ago. Support is obtained through the recalculated uranium series with electron spin resonance date of 286 ± 32 thousand years ago for a tooth from the Irhoud 3 hominin mandible. These ages are also consistent with the faunal and microfaunal assemblages and almost double the previous age estimates for the lower part of the deposits. The north African site of Jebel Irhoud contains one of the earliest directly dated Middle Stone Age assemblages, and its associated human remains are the oldest reported for H. sapiens. The emergence of our species and of the Middle Stone Age appear to be close in time, and these data suggest a larger scale, potentially pan-African, origin for both.

Pleistocene Hominins as a Resource for Carnivores: A c. 500,000-Year-Old Human Femur Bearing Tooth-Marks in North Africa (Thomas Quarry I, Morocco).

In many Middle Pleistocene sites, the co-occurrence of hominins with carnivores, who both contributed to faunal accumulations, suggests competition for resources as well as for living spaces. Despite this, there is very little evidence of direct interaction between them to-date. Recently, a human femoral diaphysis has been recognized in South-West of Casablanca (Morocco), in the locality called Thomas Quarry I. This site is famous for its Middle Pleistocene fossil hominins considered representatives of Homo rhodesiensis. The bone was discovered in Unit 4 of the Grotte à Hominidés (GH), dated to c. 500 ky and was associated with Acheulean artefacts and a rich mammalian fauna. Anatomically, it fits well within the group of known early Middle Pleistocene Homo, but its chief point of interest is that the diaphyseal ends display numerous tooth marks showing that it had been consumed shortly after death by a large carnivore, probably a hyena. This bone represents the first evidence of consumption of human remains by carnivores in the cave. Whether predated or scavenged, this chewed femur indicates that humans were a resource for carnivores, underlining their close relationships during the Middle Pleistocene in Atlantic Morocco.

Middle Pleistocene ecology and Neanderthal subsistence: insights from stable isotope analyses in Payre (Ardèche, southeastern France).

The Middle Palaeolithic site of Payre in southeastern France yields abundant archaeological material associated with fossil hominid remains. With its long sequence of Middle Pleistocene deposits, Payre is a key site to study the Middle Palaeolithic chronology of this region. This study is the first to investigate carbon and oxygen isotope contents of Neanderthal tooth enamel bioapatite, together with a wide range of herbivorous and carnivorous species. The aim is to contribute to the understanding of hunting behaviour, resource partitioning, diet and habitat use of animals and Neanderthals through a palaeoecological reconstruction. Local topography had a visible influence on carbon and oxygen stable isotope values recorded in herbivore tooth enamel. This was used to investigate possible habitats of herbivores. The different herbivorous species do not show large variations of their carbon and oxygen isotope values through time, indicating niche conservatism from OIS 8-7 to OIS 6-5, i.e., independently of palaeoclimatic and palaeoenvironmental variations. Based on these new observations, we conclude that Neanderthals employed a stable subsistence strategy over time, using a variety of local resources, with resource partitioning visible between humans and carnivores, especially wolves. A comparison of the results of stable isotopic investigation with the results of tooth wear analyses previously conducted on the same teeth allowed us to demonstrate that grazing and browsing do not bind animals to a specific habitat in a C3 environment as reflected in the isotopic values.

A chronological framework for a long and persistent archaeological record: Melka Kunture, Ethiopia.

New (40)Ar/(39)Ar geochronological data for several volcanic ash horizons from Melka Kunture, Ethiopia, allow for significantly more precise age constraints to be placed upon the lithostratigraphy, archaeology and paleontology from this long record. Ashes from the Melka Kunture Formation at Gombore yielded the most reliable age constraints, from 1.393 ± 0.162 Ma(2) (millions of years ago) near the base of the section to 0.709 ± 0.013 Ma near the top. Dating the Garba section proved more problematic, but the base of the section, which contains numerous Oldowan obsidian artifacts, may be >1.719 ± 0.199 Ma, while the top is securely dated to 0.869 ± 0.020 Ma. The large ignimbrite from the Kella Formation at Kella and Melka Garba is dated to 1.262 ± 0.034 Ma and pre-dates Acheulean artifacts in the area. The Gombore II site, which has yielded two Homo skull fragments, 'twisted bifaces,' and a preserved butchery site, is now constrained between 0.875 ± 0.010 Ma and 0.709 ± 0.013 Ma. Additional ashes from these and other sites further constrain the timing of deposition throughout the section. Integration with previously published magnetostratigraphy has allowed for the first time a relatively complete, reliable timeline for the deposition of sediments, environmental changes, archaeology, and paleontology at Melka Kunture.